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4D Berry White - Daisybushes - African Daisies- African Daisy - South African Daisy - Cape Daisy - Blue-Eyed Daisy (Osteospermum) by [email protected]

© [email protected], all rights reserved.

4D Berry White - Daisybushes - African Daisies- African Daisy - South African Daisy - Cape Daisy - Blue-Eyed Daisy (Osteospermum)

Osteospermum is a genus of flowering plants belonging to the Calenduleae, one of the smaller tribes of the sunflower/daisy family Asteraceae. They are known as the daisybushes or African daisies. Its species have been given several common names, including African daisy, South African daisy, Cape daisy and blue-eyed daisy.

Their alternate (rarely opposite) leaves are green, but some variegated forms exist. The leaf form is lanceolate. The leaf margin is entire, but hardy types are toothed.

The daisy-like composite flower consists of disc florets and ray florets, growing singly at the end of branches or sometimes in inflorescences of terminal corymbose cymes. The disc florets are pseudo-bisexual and come in several colors such as blue, yellow and purple. The hardy types usually show a dark blue center in the disc until the yellow pollen is shed. The ray florets are female and are found diverse colors such as white, cream, pink, purple, mauve to yellow. Some cultivars have "spooned" petals such as "Pink Whirls". Many species flower a second time late summer, stimulated by the cooler night temperatures. Hardy types show profuse flowering in the spring, but they do not get a second flush of flowers.

The genus Osteospermum was named by Carl Linnaeus in 1753. The scientific name is derived from the Greek osteon (bone) and Latin spermum (seed). Plants of the World Online and the South African National Biodiversity Institute accept a broad definition of the genus, which is a sister taxon to Dimorphotheca. Other authorities treat its generic synonyms (Chrysanthemoides, Oligocarpus, etc.) as separate genera.

74 species are accepted:
Osteospermum acanthospermum (DC.) Norl.
Osteospermum aciphyllum DC.
Osteospermum afromontanum Norl.
Osteospermum amplectens (Harv.) Norl.
Osteospermum angolense Norl.
Osteospermum apterum (B.Nord.) J.C.Manning & Goldblatt
Osteospermum armatum Norl.
Osteospermum asperulum (DC.) Norl.
Osteospermum attenuatum Hilliard & B.L.Burtt
Osteospermum auriculatum (S.Moore) Norl.
Osteospermum australe B.Nord.
Osteospermum bidens Thunb.
Osteospermum bolusii (Compton) Norl.
Osteospermum breviradiatum Norl. — Lemoenboegoe
Osteospermum burttianum B.Nord.
Osteospermum calcicola (J.C.Manning & Goldblatt) J.C.Manning & Goldblatt
Osteospermum calendulaceum L.f. — Stinking Roger
Osteospermum ciliatum P.J.Bergius
Osteospermum connatum DC.
Osteospermum corymbosum L.
Osteospermum crassifolium (O.Hoffm.) Norl.
Osteospermum dentatum Burm.f.
Osteospermum elsieae Norl.
Osteospermum grandidentatum DC. — Yellow trailing daisy
Osteospermum grandiflorum DC.
Osteospermum hafstroemii Norl.
Osteospermum herbaceum L.f.
Osteospermum hirsutum Thunb.
Osteospermum hispidum Harv.
Osteospermum hyoseroides (DC.) Norl.
Osteospermum ilicifolium L.
Osteospermum imbricatum L.
Osteospermum incanum Burm.f.
Osteospermum junceum P.J.Bergius
Osteospermum karrooicum (Bolus) Norl.
Osteospermum lanceolatum DC.
Osteospermum leptolobum (Harv.) Norl.
Osteospermum microcarpum (Harv.) Norl.
Osteospermum microphyllum DC.
Osteospermum moniliferum L.
Osteospermum monocephalum (Oliv. & Hiern) Norl.
Osteospermum monstrosum (Burm.f.) J.C.Manning & Goldblatt
Osteospermum montanum Klatt
Osteospermum muricatum E.Mey. ex DC.
Osteospermum namibense Swanepoel
Osteospermum nervosum (Hutch.) Norl.
Osteospermum nordenstamii J.C.Manning & Goldblatt
Osteospermum norlindhianum J.C.Manning & Goldblatt
Osteospermum nyikense Norl.
Osteospermum oppositifolium (Aiton) Norl.
Osteospermum pinnatilobatum Norl.
Osteospermum pinnatum (Thunb.) Norl.
Osteospermum polycephalum (DC.) Norl.
Osteospermum polygaloides L.
Osteospermum potbergense A.R.Wood & B.Nord.
Osteospermum pterigoideum Klatt
Osteospermum pyrifolium Norl.
Osteospermum rigidum Aiton
Osteospermum rosulatum Norl.
Osteospermum rotundifolium (DC.) Norl.
Osteospermum sanctae-helenae Norl.
Osteospermum scariosum DC.
Osteospermum sinuatum (DC.) Norl.
Osteospermum spathulatum (DC.) Norl.
Osteospermum spinescens Thunb.
Osteospermum spinigerum (Norl.) Norl.
Osteospermum spinosum L.
Osteospermum striatum Burtt Davy
Osteospermum subulatum DC.
Osteospermum thodei Markötter
Osteospermum tomentosum (L.f.) B.Nord.
Osteospermum triquetrum L.f.
Osteospermum vaillantii (Decne.) Norl.
Osteospermum volkensii (O.Hoffm.) Norl.
Formerly placed here
Dimorphotheca barberae — synonym Osteospermum barberae (Harv.) Norl.)
Dimorphotheca ecklonis — Cape marguerite, blue-and-white daisybush (synonym Osteospermum ecklonis (DC.) Norl.)
Dimorphotheca fruticosa — Trailing African daisy, shrubby daisybush (synonym Osteospermum fruticosum (L.) Norl.)
Dimorphotheca jucunda (E.Phillips) Norl. — South African daisy (synonym Osteospermum jucundum E.Phillips)

There are about 70 species native to southern and eastern Africa and the Arabian peninsula.

Osteospermum are popular in cultivation, where they are frequently used in summer bedding schemes in parks and gardens. Numerous hybrids and cultivars have been grown with a wide range of tropical colors. Yellow cultivars tend to have a yellow center (sometimes off-white).

Plants prefer a warm and sunny position and rich soil, although they tolerate poor soil, salt or drought well. Modern cultivars flower continuously when watered and fertilised well, and dead-heading is not necessary, because they do not set seed easily. If planted in a container, soil should be prevented from drying out completely. If they do, the plants will go into "sleep mode" and survive the period of drought, but they will abort their flower buds and not easily come back into flower. Moreover, roots are relatively susceptible to rotting if watered too profusely after the dry period.

Most widely sold cultivars are grown as annuals, are mainly hybrids of O. jucundum, O. ecklonis and O. grandiflorum and can be hardy to -2 °C (30 °F). If hardy, they can be grown as perennials or as shrubs.

Cultivars (those marked agm have gained the Royal Horticultural Society's Award of Garden Merit):
'White Spoon'
'Acapulco'
'African Queen'
'Apricot'
'Biera'
'Big Pink'
‘Blackthorn Seedling’ agm
'Bodegas Pink'
'Buttermilk' agm
'Chris Brickell'
'Duet'
'Giles Gilbey'
'Hopleys’ agm
'Ice White'
'Langtrees agm
’Lady Leitrim’ agm
'Lilac Spoon'
'Marbella'
'Merriments Joy'
'Nairobi Purple'
O. jucundum agm
'Passion Mix'
'Pink'
'Pink Beauty'
'Pink Whirls' agm
'Silver Sparkler' agm
'Soprano'
'Starshine'
'Springstar Gemma'
'Sunkist'
'Weetwood'
'White Pim' agm
'White Spoon'
'White Whirls'

Marigold (Tagetes) by [email protected]

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Marigold (Tagetes)

Tagetes is a genus of 50 species of annual or perennial, mostly herbaceous plants in the family Asteraceae. They are among several groups of plants known in English as marigolds. The genus Tagetes was described by Carl Linnaeus in 1753.

These plants are native to Mexico, growing naturally from Mexico's valley down to the south and even reaching several other Latinamerican countries, but some species have become naturalized around the world. One species, T. minuta, is considered a noxious invasive plant in some areas.

Description
Tagetes species vary in size from 0.1 to 2.2 m tall. Most species have pinnate green leaves. Blooms naturally occur in golden, orange, yellow, and white colors, often with maroon highlights. Floral heads are typically (1-) to 4–6 cm diameter, generally with both ray florets and disc florets. In horticulture, they tend to be planted as annuals, although the perennial species are gaining popularity. They have fibrous roots.

Depending on the species, Tagetes species grow well in almost any sort of soil. Most horticultural selections grow best in soil with good drainage, and some cultivars are known to have good tolerance to drought.

Nomenclature
The Latin Tagētes derives from the Tages in Etruscan mythology, born from plowing the earth. It likely refers to the ease with which plants of this genus come out each year either by the seeds produced in the previous year, or by the stems which regrow from the stump already in place.

The common name in English, marigold, is derived from Mary's gold, a name first applied to a similar plant native to Europe, Calendula officinalis.

The most commonly cultivated varieties of Tagetes are known variously as African marigolds (usually referring to cultivars and hybrids of Tagetes erecta), or French marigolds (usually referring to hybrids and cultivars of Tagetes patula, many of which were developed in France). The so-called signet marigolds are hybrids derived mostly from Tagetes tenuifolia.

Cultivation and uses
Depending on the species, marigold foliage has a musky, pungent scent, though some varieties have been bred to be scentless. Due to antibacterial thiophenes exuded by the roots, Tagetes should not be planted near any legume crop. Some of the perennial species are deer-, rabbit-, rodent- and javelina or peccary-resistant.

T. minuta (khakibush or huacatay), originally from South America, has been used as a source of essential oil for the perfume and industry known as tagette or "marigold oil", and as a flavourant in the food and tobacco industries. It is commonly cultivated in South Africa, where the species is also a useful pioneer plant in the reclamation of disturbed land.

The florets of Tagetes erecta are rich in the orange-yellow carotenoid lutein and are used as a food colour (INS number E161b) in the European Union for foods such as pasta, vegetable oil, margarine, mayonnaise, salad dressing, baked goods, confectionery, dairy products, ice cream, yogurt, citrus juice and mustard. In the United States, however, the powders and extracts are only approved as colorants in poultry feed.

Marigolds are recorded as a food plant for some Lepidoptera caterpillars including the dot moth, and a nectar source for other butterflies and bumblebees. They are often part of butterfly gardening plantings. In the wild, many species are pollinated by beetles.

Cultural significance
Tagetes lucida
The species Tagetes lucida, known as pericón, is used to prepare a sweetish, anise-flavored medicinal tea in Mexico. It is also used as a culinary herb in many warm climates, as a substitute for tarragon, and offered in the nursery as "Texas tarragon" or "Mexican mint marigold".

Tagetes minuta
Tagetes minuta, native to southern South America, is a tall, upright marigold plant with small flowers used as a culinary herb in Peru, Ecuador, and parts of Chile and Bolivia, where it is called by the Incan term huacatay. The paste is used to make the popular potato dish called ocopa. Having both "green" and "yellow/orange" notes, the taste and odor of fresh T. minuta is like a mixture of sweet basil, tarragon, mint and citrus. It is also used as a medicinal tea for gastrointestinal complaints and specifically against nematodes.

Tagetes erecta
Tagetes erecta is widely used in Day of the Dead celebrations in Mexico.

In Bangladesh, India and other South Asian countries, marigold is used for ornamentation purposes in functions like turmeric ceremony, weddings, Pohela Falgun and other functions. During the colonial period the native varieties of these flowers were replaced by American species like T. erecta, T. patula and T. tenuifolia. The marigold is also widely cultivated in India and Thailand, particularly the species T. erecta, Tagetes patula and T. tenuifolia. It is always sold in the markets for daily worship and rituals. Vast quantities of marigolds are used in garlands and decoration for weddings, festivals, and religious events. Marigold cultivation is extensively seen in Telangana, Andhra Pradesh, Tamil Nadu, West Bengal, Karnataka and Uttar Pradesh (for the Vijayadashami and Diwali markets) states of India.

In Ukraine, chornobryvtsi (T. erecta, T. patula and the signet marigold, l. tenuifolia) are regarded as one of the national symbols, and are often mentioned in songs, poems and tales.

Accepted species
Tagetes apetala
Tagetes arenicola
Tagetes argentina Cabrera
Tagetes biflora Cabrera
Tagetes campanulata Griseb.
Tagetes daucoides
Tagetes elliptica Sm.
Tagetes elongata
Tagetes epapposa
Tagetes erecta L. – African marigold, Aztec marigold
Tagetes filifolia Lag.
Tagetes foeniculacea
Tagetes foetidissima DC.
Tagetes hartwegii Greenm.
Tagetes iltisiana Rydb.
Tagetes inclusa
Tagetes lacera
Tagetes laxa Cabrera
Tagetes lemmonii A.Gray – Mt. Lemmon marigold, Mexican marigold[17]
Tagetes linifolia
Tagetes lucida Cav. – Mexican mint marigold, Texas tarragon
Tagetes mandonii
Tagetes mendocina Phil.
Tagetes micrantha Cav. – licorice marigold
Tagetes microglossa
Tagetes minima
Tagetes minuta L. – wild marigold
Tagetes moorei
Tagetes mulleri S.F.Blake
Tagetes multiflora Kunth
Tagetes nelsonii Greenm.
Tagetes oaxacana
Tagetes osteni
Tagetes palmeri
Tagetes parryi A.Gray
Tagetes patula L. – French marigold
Tagetes perezii Cabrera
Tagetes praetermissa
Tagetes pringlei S.Watson
Tagetes pusilla
Tagetes riojana M.Ferraro
Tagetes rupestris Cabrera
Tagetes stenophylla B.L.Rob.
Tagetes subulata Cerv.
Tagetes subvillosa
Tagetes tenuifolia Cav. – signet marigold
Tagetes terniflora Kunth
Tagetes triradiata
Tagetes verticillata Lag. & Rodr.
Tagetes zypaquirensis Humb. & Bonpl.

Sunflower (Helianthus) by [email protected]

© [email protected], all rights reserved.

Sunflower (Helianthus)

Helianthus is a genus comprising about 70 species of annual and perennial flowering plants in the daisy family Asteraceae commonly known as sunflowers. Except for three South American species, the species of Helianthus are native to North America and Central America. The best-known species is the common sunflower (Helianthus annuus). This and other species, notably Jerusalem artichoke (H. tuberosus), are cultivated in temperate regions and some tropical regions, as food crops for humans, cattle, and poultry, and as ornamental plants. The species H. annuus typically grows during the summer and into early fall, with the peak growth season being mid-summer.
Several perennial Helianthus species are grown in gardens, but have a tendency to spread rapidly and can become aggressive. On the other hand, the whorled sunflower, Helianthus verticillatus, was listed as an endangered species in 2014 when the U.S. Fish and Wildlife Service issued a final rule protecting it under the Endangered Species Act. The primary threats are industrial forestry and pine plantations in Alabama, Georgia, and Tennessee. They grow to 1.8 metres (6 feet) and are primarily found in woodlands, adjacent to creeks and moist, prairie-like areas.

The common sunflower is the national flower of Ukraine, cultivated there for several centuries.

Description

The disk of a sunflower is made up of many little flowers. The ray flowers here are dried
A field of sunflowers in North Carolina
In North Carolina

Sunflowers are usually tall annual or perennial plants that in some species can grow to a height of 300 centimetres (120 inches) or more. Each "flower" is actually a disc made up of tiny flowers, to form a larger false flower to better attract pollinators. The plants bear one or more wide, terminal capitula (flower heads made up of many tiny flowers), with bright yellow ray florets (mini flowers inside a flower head) at the outside and yellow or maroon (also known as a brown/red) disc florets inside. Several ornamental cultivars of H. annuus have red-colored ray florets; all of them stem from a single original mutant. While the majority of sunflowers are yellow, there are branching varieties in other colors including, orange, red and purple.

The petiolate leaves are dentate and often sticky. The lower leaves are opposite, ovate, or often heart-shaped. The rough and hairy stem is branched in the upper part in wild plants, but is usually unbranched in domesticated cultivars.

This genus is distinguished technically by the fact that the ray florets (when present) are sterile, and by the presence on the disk flowers of a pappus that is of two awn-like scales that are caducous (that is, easily detached and falling at maturity). Some species also have additional shorter scales in the pappus, and one species lacks a pappus entirely. Another technical feature that distinguishes the genus more reliably, but requires a microscope to see, is the presence of a prominent, multicellular appendage at the apex of the style. Further, the florets of a sunflower are arranged in a natural spiral.

Variability is seen among the perennial species that make up the bulk of those in the genus. Some have most or all of the large leaves in a rosette at the base of the plant and produce a flowering stem that has leaves that are reduced in size. Most of the perennials have disk flowers that are entirely yellow, but a few have disk flowers with reddish lobes. One species, H. radula, lacks ray flowers altogether.

Overall, the macroevolution of the Helianthus is driven by multiple biotic and abiotic factors and influences various floral morphology.

Helianthus species are used as food plants by the larvae of many lepidopterans.

Growth stages
The growth of a sunflower depends strictly on its genetic makeup and background. Additionally, the season it is planted will have effects on its development; those seasons tend to be in the middle of summer and beginning of fall. Sunflower development is classified by a series of vegetative stages and reproductive stages that can be determined by identifying the heads or main branch of a single head or branched head.

Facing the Sun (heliotropism)
Before blooming, Helianthus plant heads tilt during the day to face the Sun. This movement is referred to as heliotropism, which continues for a short time when flower buds form and young Helianthus heads track the Sun. At night, the flower heads reorient their position and face East in anticipation for the sunrise. Sunflowers move back to their original position between 3am and 6am, and the leaves follow about an hour later.

By the time they are mature and reach anthesis, Helianthus generally stop moving and remain facing east, which lets them be warmed by the rising sun. Historically, this has led to controversy on whether or not Helianthus is heliotropic, as many scientists have failed to observe movement when studying plants that have already bloomed.

This is notably different from heliotropism in leaves, as the moving mechanism for leaves exists in the pulvinus. Since flowers do not have pulvini, the movement is caused by increased growth rate of the stems. The growth rate accumulation of the stem on the east side of the stem gradually pushes the flower from east to west during daytime. This matches with the Sun as it rises from the east and falls in the west. At night, the growth rate is higher in the west side of the stem that gradually pushes the flower from the west side back to the east side. In addition, it is not actually the whole plant that changes its direction to face the Sun, but the flower itself that bends to be illuminated by the Suns rays.

The heliotropic movement is caused by growth on the opposite side of the flower, driven by accumulation of growth hormones during Sun exposure.

Heliotropism persists on cloudy days when the sun is not shining brightly, meaning that the movement is endogenous as a trained and continuous process. However, flower movement does not occur during long periods of rain or clouds. It also does not occur in a growth chamber when exposed to 16 hours of light or in greenhouses, suggesting that the plants require a directional, moving light source. Helianthus can also discriminate between different types of light. When exposed to different light frequencies, the hypocotyls will bend toward blue light but not red light, depending on the quality of the light source.

It is the circadian rhythms and the differences of the stem growth rate that work together and cause the heliotropism of the Helianthus. This is important for attracting pollinators and increasing growth metabolism. Future studies are needed for identifying the exact physiological basis and cellular mechanism for this behavior.

Taxonomy
Helianthus is derived from Greek ἥλιος hēlios "sun" and ἄνθος ánthos "flower", because its round flower heads in combination with the ligules look like the Sun.

There are many species recognized in the genus:

Helianthus agrestis Pollard – southeastern sunflower – Florida, Georgia
Helianthus ambiguus Britt. – Wisconsin, Michigan, Ohio, New York
Helianthus angustifolius L. – swamp sunflower – Texas, northern Florida to southern Illinois, Long Island, New York
Helianthus annuus L. – common sunflower, girasol – most of United States + Canada
Helianthus anomalus S.F.Blake – western sunflower – Nevada, Utah, Arizona, New Mexico
Helianthus argophyllus Torr. & A.Gray – silverleaf sunflower – Texas, North Carolina, Florida
Helianthus arizonensis R.C.Jacks. – Arizona sunflower – Arizona, New Mexico
Helianthus atrorubens L. – purpledisk sunflower – Louisiana, Alabama, Georgia, Florida, South Carolina, North Carolina, Tennessee, Kentucky, Virginia
Helianthus bolanderi A.Gray – serpentine sunflower – California, Oregon
Helianthus × brevifolius E.Watson – Texas, Indiana, Ohio
Helianthus californicus DC. – California sunflower – California
Helianthus carnosus Small – lakeside sunflower – Florida
Helianthus ciliaris DC. – Texas blueweed – United States: Washington, California, Arizona, New Mexico, Nevada, Utah, Texas, Oklahoma, Colorado, Kansas, Illinois; Mexico: Tamaulipas, Coahuila, Chihuahua, Sonora
Helianthus cinereus Small – Missouri, Kentucky, Indiana, Ohio
Helianthus coloradensis Cockerell – prairie sunflower – Colorado, New Mexico
Helianthus cusickii A.Gray – Cusick's sunflower – Washington, Oregon, California, Idaho, Nevada
Helianthus debilis Nutt. – cucumberleaf sunflower – Texas to Maine, Mississippi
Helianthus decapetalus L. – thinleaf sunflower – eastern United States; Ontario, Quebec
Helianthus deserticola Heiser – desert sunflower – Arizona, Nevada, Utah
†Helianthus diffusus Sims – Missouri†
Helianthus dissectifolius R.C.Jacks. – Mexico
Helianthus divaricatus L. – woodland sunflower or rough woodland sunflower – eastern United States; Ontario, Quebec
Helianthus × divariserratus R.W.Long Michigan, Indiana, Ohio, Connecticut
Helianthus × doronicoides Lam. – Texas, Oklahoma, Arkansas, Missouri, Iowa, Minnesota, Illinois, Kentucky, Indiana, Ohio, Pennsylvania, Michigan, New Jersey, Virginia
Helianthus eggertii Small – Alabama, Kentucky, and Tennessee
Helianthus exilis A.Gray – California
Helianthus floridanus A.Gray ex Chapm. – Florida sunflower – Louisiana, Alabama, Georgia, Florida, South Carolina, North Carolina
Helianthus giganteus L. – giant sunflower – eastern United States; most of Canada
Helianthus glaucophyllus D.M.Sm – whiteleaf sunflower – Tennessee, South Carolina, North Carolina
Helianthus × glaucus Small – scattered locales in southeastern United States
Helianthus gracilentus A.Gray – slender sunflower – California
Helianthus grosseserratus M.Martens – sawtooth sunflower – Great Plains, Great Lakes, Ontario, Quebec
Helianthus heterophyllus Nutt. – variableleaf sunflower – Coastal plain of Texas to North Carolina
Helianthus hirsutus Raf. – hairy sunflower – central and eastern United States, Ontario
Helianthus × intermedius R.W.Long – intermediate sunflower – scattered locales in United States
Helianthus laciniatus A.Gray – alkali sunflower – United States: Arizona, New Mexico, Texas; Mexico: Coahuila, Nuevo León
Helianthus × laetiflorus Pers. – cheerful sunflower, mountain sunflower – scattered in eastern and central United States; Canada
Helianthus laevigatus Torr. & A.Gray – smooth sunflower – Georgia, South Carolina, North Carolina, Virginia, Maryland, West Virginia
Helianthus lenticularis Douglas ex Lindl. California, Texas
Helianthus longifolius Pursh – longleaf sunflower – Alabama, Georgia, North Carolina
Helianthus × luxurians (E.Watson) E.Watson – Great Lakes region
Helianthus maximiliani Schrad. – Maximillian sunflower – much of United States and Canada
Helianthus membranifolius Poir. – French Guiana
Helianthus microcephalus Torr. & A.Gray – eastern United States
Helianthus mollis Lam. – downy sunflower, ashy sunflower – Ontario, eastern and central United States
Helianthus multiflorus L. – manyflower sunflower – Ohio
Helianthus navarri Phil. – Chile
Helianthus neglectus Heiser – neglected sunflower – New Mexico, Texas
Helianthus niveus (Benth.) Brandegee – showy sunflower – United States: California, Arizona; Mexico: Baja California, Baja California Sur
Helianthus nuttallii Torr. & A.Gray – western and central United States, Canada
Helianthus occidentalis Riddell – fewleaf sunflower, western sunflower – Great Lakes region, scattered in southeastern United States
Helianthus × orgyaloides Cockerell – Colorado, Kansas
Helianthus paradoxus Heiser – paradox sunflower – Utah, New Mexico, Texas
Helianthus pauciflorus Nutt. – stiff sunflower – central United States, Canada
Helianthus petiolaris Nutt. – prairie sunflower, lesser sunflower – much of United States, Canada
Helianthus porteri (A.Gray) Pruski – Porter's sunflower – Alabama, Georgia, South Carolina, North Carolina
Helianthus praecox Engelm. & A.Gray Texas sunflower – Texas
†Helianthus praetermissus – New Mexico sunflower – New Mexico†
Helianthus pumilus Nutt. – little sunflower – Colorado, Wyoming, Montana, Utah, Idaho
Helianthus radula (Pursh) Torr. & A.Gray – rayless sunflower – Louisiana, Mississippi, Alabama, Georgia, South Carolina, Florida
Helianthus resinosus Small – resindot sunflower – Mississippi, Alabama, Georgia, South Carolina, North Carolina, Florida
Helianthus salicifolius A.Dietr. – willowleaf sunflower – Texas, Oklahoma, Kansas, Missouri, Illinois, Wisconsin, Ohio, Pennsylvania, New York
Helianthus sarmentosus Rich. – French Guiana
Helianthus scaberrimus Elliott – South Carolina
Helianthus schweinitzii Torr. & A.Gray – Schweinitz's sunflower – South Carolina, North Carolina
Helianthus silphioides Nutt. – rosinweed sunflower – Lower Mississippi Valley
Helianthus simulans E.Watson – muck sunflower – southeastern United States
Helianthus smithii Heiser – Smith's sunflower – Alabama, Georgia, Tennessee
Helianthus speciosus Hook. – Michoacán
Helianthus strumosus L. – eastern and central United States, Canada
Helianthus subcanescens (A.Gray) E.Watson – Manitoba, north-central United States
Helianthus subtuberosus Bourg.
Helianthus tuberosus L. – Jerusalem artichoke, sunchoke, earth-apple, topinambur – much of United States and Canada
Helianthus verticillatus Small – whorled sunflower – Alabama, Georgia, Tennessee
Formerly included
The following species were previously included in the genus Helianthus.

Flourensia thurifera (Molina) DC. (as H. thurifer Molina)
Helianthella quinquenervis (Hook.) A.Gray (as H. quinquenervis Hook.)
Helianthella uniflora (Nutt.) Torr. & A.Gray (as H. uniflorus Nutt.)
Pappobolus imbaburensis (Hieron.) Panero (as H. imbaburensis Hieron.)
Viguiera procumbens (Pers.) S.F.Blake (as H. procumbens Pers.)
Uses
The seeds of H. annuus are used as human food. Most cultivars of sunflower are variants of H. annuus, but four other species (all perennials) are also domesticated. This includes H. tuberosus, the Jerusalem artichoke, which produces edible tubers.

There are many species in the sunflower genus Helianthus, and many species in other genera that may be called sunflowers.

The Maximillian sunflower (Helianthus maximiliani) is one of 38 species of perennial sunflower native to North America. The Land Institute and other breeding programs are currently exploring the potential for these as a perennial seed crop.
The sunchoke (Jerusalem artichoke or Helianthus tuberosus) is related to the sunflower, another example of perennial sunflower.
The Mexican sunflower is Tithonia rotundifolia. It is only very distantly related to North American sunflowers.
False sunflower refers to plants of the genus Heliopsis.
Ecology

Sunflowers have been proven to be excellent plants to attract beneficial insects, including pollinators. Helianthus spp. are a nectar producing flowering plant that attract pollinators and parasitoids which reduce the pest populations in nearby crop vegetation. Sunflowers attract different beneficial pollinators (e.g., honey bees) and other known insect prey to feed on and control the population of parasitic pests that could be harmful to the crops. Predacious insects are first attracted to sunflowers once they are planted. Once the Helianthus spp. reaches six inches and produces flowers it begins to attract more pollinators. Distance between sunflower rows and crop vegetation plays an important role in this phenomenon, hypothesizing that closer proximity to the crops will increase insect attraction.

In addition to pollinators of Helianthus spp., there are other factors such as abiotic stress, florivory, and disease which also contribute to the evolution of floral traits. These selective pressures, which stem from several biotic and abiotic factors are associated with habitat environmental conditions which all play a role in the overall morphology of the sunflowers' floral traits.

An ecosystem is composed of both biotic (which are living elements of an ecosystem such as plants, animals, fungi, protists, and bacteria), and abiotic factors (non-living elements of an ecosystem such as air, soil, water, light, salinity and temperature).

It is thought that two biotic factors can explain for the evolution of larger sunflowers and why they are present in more drier environments. For one thing, the selection by pollinators is thought to have increased the sunflower's size in a drier environment. This is because in a drier environment, there are typically less pollinators. As a result, in order for the sunflower to be able to attract more pollinators, they had to increase the morphology of their floral traits in that they had to increase their display size. Another biotic factor that can explain for the evolution of larger sunflowers in drier environments is that the pressure from florivory and disease favors smaller flowers in habitats that have a more moderate supply of moisture (mesic habitat). Wetter environments usually have more dense vegetation, more herbivores, and more surrounding pathogens. As larger flowers are typically more susceptible to disease and florivory, smaller flowers may have evolved in wetter environments which explains the evolution of larger sunflowers in more drier environments.

China Aster - Annual Aster (Callistephus) by [email protected]

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China Aster - Annual Aster (Callistephus)

Callistephus is a monotypic genus of flowering plants in the aster family, Asteraceae, containing the single species Callistephus chinensis. Its common names include China aster and annual aster. It is native to China and Korea. and it is cultivated worldwide as an ornamental plant in cottage gardens and as a cut flower.

Description
This is an annual or biennial plant with one erect, mostly unbranched stem growing 20–100 cm (7.9–39.4 in) tall. The alternately arranged leaves vary in shape or size. The basal leaves usually fall away before flowering. Leaves around the middle of the stem are a few centimeters long and are borne on winged petioles. The blades have serrated edges. Leaves toward the top may have smooth edges. The large solitary flower head grows at the top of the stem and sometimes on branches. The head is lined with layers of phyllaries, those in the outer layer large and leaflike, measuring up to 3 centimeters long. The inner layer can be purple-tinged. The head contains one or two rings of ray florets, most often in shades of reddish purple. The flowering period is from August to October in the northern hemisphere.

Cultivation
This species has been grown and bred extensively in cultivation. Cultivars are available with flowers of many colors, varying heights, and single and double heads. In the wild, they are purple in color. There are several cultivars with flowers of different colors. Cultivars with flowers of other colors were obtained in particular by hybridization. Cultivated varieties can have ray florets in most any color, including red, pink, blue, purple, and white. They are up to 3.5 centimeters long. There are many yellow disc florets in the center. The fruit is a rough-textured, glandular, purple-mottled cypsela that turns gray with age. It has a pappus composed of one outer layer of reddish scales and two inner layers of white bristles.

It has been in cultivation in Europe since 1728 at the latest. In China, it has been cultivated for 2000 years.

Distribution
This plant is a naturalized species in some areas outside of its native range (Indochina, Europe, North America, Australia, New Zealand, etc.) and can be found on the edges of deciduous forests. It grows at altitudes from 300 to 2700 above mean sea level. As ornamental plant it thrives on fresh, nutrient-rich, easy-humus rich soils in warm humid air conditioning.

Pests and diseases
The plants are susceptible to fusarium wilt, a fungal disease. Some cultivars are more resistant to this than others. Pest insects can include leafhoppers, aphids, blister beetles, and the tarnished plant bug. The plant is also susceptible to mites.[

Dahlia (Asteraceae) by [email protected]

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Dahlia (Asteraceae)

Dahlia is a genus of bushy, tuberous, herbaceous perennial plants native to Mexico and Central America. As a member of the Asteraceae (former name: Compositae) family of dicotyledonous plants, its relatives include the sunflower, daisy, chrysanthemum, and zinnia. There are 49 species of dahlia, with flowers in almost every hue (except blue), with hybrids commonly grown as garden plants.

Dahlias were known to the Aztecs until their Spanish conquest, after which the plants were brought to Europe. The (high in sugar) tubers of some varieties are of value to humans.

Description


Complete Stages of Dahlia Flower: (start from 2nd row 1st pic clockwise) Complete Bloomed Flower, Blooming Bud. Bud, Bud in The tree And Dahlia Plant; Captured In NIT Agartala, India
Dahlias are perennial plants with tuberous roots, though they are grown as annuals in some regions with cold winters. While some have herbaceous stems, others have stems which lignify in the absence of secondary tissue and resprout following winter dormancy, allowing further seasons of growth. As a member of the Asteraceae, the dahlia has a flower head that is actually a composite (hence the older name Compositae) with both central disc florets and surrounding ray florets. Each floret is a flower in its own right, but is often incorrectly described as a petal. The modern name Asteraceae refers to the appearance of a star with surrounding rays.

The stems are leafy, ranging in height from as low as 30 centimetres (12 inches) to more than 1.8–2.4 metres (6–8 feet). Flower forms are variable, with one head per stem; these can be as small as 5 cm (2 in) in diameter or up to 30 cm (1 ft) ("dinner plate"). The majority of species do not produce scented flowers. Like most plants that do not attract pollinating insects through scent, they are brightly colored, displaying most hues, with the exception of blue.

The great variety in species results from garden dahlias being octoploids, having eight sets of homologous chromosomes. In addition, dahlias also contain many transposons—genetic pieces that move from place to place upon an allele—which contributes to their manifesting such great diversity.

Taxonomy
Taxonomic history
Early history
Dahlia
Dahlia
Orange Dahlia
Orange Dahlia
Spaniards reported finding the plants growing in Mexico in 1525, but the earliest known description is by Francisco Hernández, physician to Philip II, who was ordered to visit Mexico in 1570 to study the "natural products of that country". They were used as a source of food by the indigenous peoples, who both gathered wild specimens and cultivated crops. The indigenous peoples variously identified the plants as "Chichipatl" (Toltecs) and "Acocotle" or "Cocoxochitl" (Aztecs). From Hernandez's perception of Nahuatl to Spanish (through various other translations) the word is "water cane", "water pipe", "water pipe flower", "hollow stem flower", or "cane flower", all referring to the hollow plant stems.

Hernandez described two varieties of dahlias (the pinwheel-like Dahlia pinnata and the huge Dahlia imperialis) as well as other medicinal plants of New Spain. Francisco Dominguez, an Hidalgo gentleman who accompanied Hernandez on part of his seven-year study, made a series of drawings to supplement the four volume report. Three of his drawings showed plants with flowers: two resembled the modern bedding dahlia, and one resembled the species Dahlia merckii; all displayed a high degree of doubleness. In 1578, a manuscript titled Nova Plantarum, Animalium et Mineralium Mexicanorum Historia, was sent back to the Escorial in Madrid. It was translated into Latin by Francisco Ximenes in 1615. In 1640, Francisco Cesi, President of the Academia dei Lincei of Rome, bought the Ximenes translation and, after annotating it, published it in 1649–1651 as two volumes, Rerum Medicarum Novae Hispaniae Thesaurus Seu Nova Plantarium, Animalium et Mineralium Mexicanorum Historia. The original manuscripts were destroyed in a fire in the mid-1600s.

European introduction
In 1787, the French botanist Nicolas-Joseph Thiéry de Menonville, sent to Mexico to steal the cochineal insect valued for its scarlet dye, reported the strangely beautiful flowers he had seen growing in a garden in Oaxaca. In 1789, Vicente Cervantes, director of the Botanical Garden at Mexico City, sent "plant parts" to Abbe Antonio José Cavanilles, director of the Royal Gardens of Madrid. Cavanilles flowered one plant that same year, then the second one a year later. In 1791 he called the new growths "Dahlia" for Anders Dahl. The first plant was called Dahlia pinnata after its pinnate foliage; the second, Dahlia rosea for its rose-purple color. In 1796, from the parts sent by Cervantes, Cavanilles flowered a third plant, which he named Dahlia coccinea for its scarlet color.

In 1798, Cavanilles sent D. pinnata seeds to Parma, Italy. That year, the Marchioness of Bute, wife of the Earl of Bute, the English Ambassador to Spain, obtained a few seeds from Cavanilles and sent them to Kew Gardens, where they flowered but were lost after two to three years.

In the following years Madrid sent seeds to Berlin and Dresden in Germany, and to Turin and Thiene in Italy. In 1802, Cavanilles sent tubers of "these three" (D. pinnata, D. rosea, D. coccinea) to Swiss botanist Augustin Pyramus de Candolle at University of Montpelier in France, Andre Thouin at the Jardin des Plantes in Paris and Scottish botanist William Aiton at Kew Gardens. That same year, John Fraser, English nurseryman and later botanical collector to the Czar of Russia, brought D. coccinea seeds from Paris to the Apothecaries Gardens in England, where they flowered in his greenhouse a year later, providing Botanical Magazine with an illustration.

In 1804, a new species, Dahlia sambucifolia, was successfully grown at Holland House, Kensington. Whilst in Madrid in 1804, Lady Holland was given either dahlia seeds or tubers by Cavanilles. She sent them back to England, to Lord Holland's librarian at Holland House, who successfully raised the plants and produced two double flowers a year later. The plants raised in 1804 did not survive; new stock was brought from France in 1815. In 1824, Lord Holland sent his wife a note containing the following verse:

The dahlia you brought to our isle
Your praises for ever shall speak;
Mid gardens as sweet as your smile,
And in colour as bright as your cheek.

In 1805, German naturalist Alexander von Humboldt sent more seeds from Mexico to Aiton in England, Thouin in Paris, and Christoph Friedrich Otto, director of the Berlin Botanical Garden. More significantly, he sent seeds to botanist Carl Ludwig Willdenow in Germany. Willdenow now reclassified the rapidly growing number of species, changing the genus from Dahlia to Georgina; after naturalist Johann Gottlieb Georgi. He combined the Cavanilles species D. pinnata and D. rosea under the name of Georgina variabilis; D. coccinea was still held to be a separate species, which he renamed Georgina coccinea.

Since 1789 when Cavanilles first flowered the dahlia in Europe, there has been an ongoing effort by many growers, botanists and taxonomists, to determine the development of the dahlia to modern times. At least 85 species have been reported: approximately 25 of these were first reported from the wild; the remainder appeared in gardens in Europe. They were considered hybrids, the results of crossing between previously reported species, or developed from the seeds sent by Humboldt from Mexico in 1805, or perhaps from some other undocumented seeds that had found their way to Europe. Several of these were soon discovered to be identical with earlier reported species, but the greatest number are new varieties. Morphological variation is highly pronounced in the dahlia. William John Cooper Lawrence, who hybridized hundreds of families of dahlias in the 1920s, stated: "I have not yet seen any two plants in the families I have raised which were not to be distinguished one from the other. Constant reclassification of the 85 reported species has resulted in a considerably smaller number of distinct species, as there is a great deal of disagreement today between systematists over classification.

In 1829, all species growing in Europe were reclassified under an all-encompassing name of D. variabilis, Desf., though this is not an accepted name. Through the interspecies cross of the Humboldt seeds and the Cavanilles species, 22 new species were reported by that year, all of which had been classified in different ways by several different taxonomists, creating considerable confusion as to which species was which. As of now Dahlias are classified into 15 different species by botanist Liberty Hyde Bailey.

In 1830 William Smith suggested that all dahlia species could be divided into two groups for color, red-tinged and purple-tinged. In investigating this idea Lawrence determined that with the exception of D. variabilis, all dahlia species may be assigned to one of two groups for flower-colour: Group I (ivory-magenta) or Group II (yellow-orange-scarlet).

Modern classification
The genus Dahlia is situated in the Asteroideae subfamily of the Asteraceae, in the Coreopsideae tribe. Within that tribe it is the second largest genus, after Coreopsis, and appears as a well defined clade within the Coreopsideae.

Subdivision
Infrageneric subdivision
See also: List of Dahlia species
Sherff (1955), in the first modern taxonomy described three sections for the 18 species he recognised, Pseudodendron, Epiphytum and Dahlia. By 1969 Sørensen recognised 29 species and four sections by splitting off Entemophyllon from section Dahlia. By contrast Giannasi (1975) using a phytochemical analysis based on flavonoids, reduced the genus to just two sections, Entemophyllon and Dahlia, the latter having three subsections, Pseudodendron, Dahlia, and Merckii. Sørensen then issued a further revision in 1980, incorporating subsection Merckii in his original section Dahlia. When he described two new species in the 1980s (Dahlia tubulata and D. congestifolia), he placed them within his existing sections. A further species, Dahlia sorensenii was added by Hansen and Hjerting in (1996). At the same time they demonstrated that Dahlia pinnata should more properly be designated D. x pinnata. D. x pinnata was shown to actually be a variant of D. sorensenii that had acquired hybrid qualities before it was introduced to Europe in the sixteenth century and formally named by Cavanilles. The original wild D. pinnata is presumed extinct. Further species continue to be described, Saar (2003) describing 35 species. However separation of the sections on morphological, cytologal and biocemical criteria has not been entirely satisfactory.

To date these sectional divisions have not been fully supported phylogenetically, which demonstrate only section Entemophyllon as a distinct sectional clade. The other major grouping is the core Dahlia clade (CDC), which includes most of the section Dahlia. The remainder of the species occupy what has been described as the variable root clade (VRC) which includes the small section Pseudodendron but also the monotypic section Epiphytum and a number of species from within section Dahlia. Outside of these three clades lie D. tubulata and D. merckii as a polytomy.

Horticulturally the sections retain some usage, section Pseudodendron being referred to as 'Tree Dahlias', Epiphytum as the 'Vine Dahlia'. The remaining two herbaceous sections being distinguished by their pinnules, opposing (Dahlia) or alternating (Entemophyllon).

Sections
Sections (including chromosome numbers), with geographical distribution;

Epiphytum Sherff (2n = 32)
10 m tall climber with aerial roots 5 cm thick and up to more than 20 m long; pinnules opposite
1 species, D. macdougallii Sherff
Mexico: Oaxaca
Entemophyllon P. D. Sorensen (2n = 34)
6 species
Mexico: Hidalgo, Nuevo León, Tamaulipas, Querétaro, Durango, San Luis Potosí
Pseudodendron P. D. Sorensen (2n = 32)
3 species + D. excelsa of uncertain identity
Mexico: Chiapas, Guerrero, Jalisco, Michoacan, Oaxaca, and Costa Rica, El Salvador, Guatemala & Colombia
Dahlia (2n = 32, 36 or 64)
24 species
Mexico: Distrito Federal, Guerrero, Hidalgo, Morelos, Nuevo León, Puebla, San Luis Potosí, Tamaulipas, Veracruz, Oaxaca, Puebla, Chiapas, México, Huehuetenango, Chihuahua, Durango, Michoacan & Guatemala
Only Pseudodendron (D. imperialis) and Dahlia (D. australis, D. coccinea) occur outside Mexico.

Species
Main article: List of Dahlia species
There are currently 42 accepted species in the genus Dahlia but new species continue to be described.

Etymology
The naming of the plant itself has long been a subject of some confusion. Many sources state that the name "Dahlia" was bestowed by the pioneering Swedish botanist and taxonomist Carl Linnaeus to honor his late student, Anders Dahl, author of Observationes Botanicae. However, Linnaeus died in 1778, more than eleven years before the plant was introduced into Europe in 1789, so while it is generally agreed that the plant was named in 1791 in honor of Dahl, who had died two years before, Linnaeus could not have been the one who did so. It was probably Abbe Antonio Jose Cavanilles, Director of the Royal Gardens of Madrid, who should be credited with the attempt to scientifically define the genus, since he not only received the first specimens from Mexico in 1789, but named the first three species that flowered from the cuttings.

Regardless of who bestowed it, the name was not so easily established. In 1805, German botanist Carl Ludwig Willdenow, asserting that the genus Dahlia Thunb. (published a year after Cavanilles's genus and now considered a synonym of Trichocladus) was more widely accepted, changed the plants' genus from Dahlia to Georgina (after the German-born naturalist Johann Gottlieb Georgi, a professor at the Imperial Academy of Sciences of St. Petersburg, Russia). He also reclassified and renamed the first three species grown, and identified, by Cavanilles. It was not until 1810, in a published article, that he officially adopted the Cavanilles's original designation of Dahlia. However, the name Georgina still persisted in Germany for the next few decades. In Russian, it is still named Georgina (Russian: георгинa).

Distribution and habitat
Dahlia is found predominantly in Mexico, but some species are found ranging as far south as northern South America. D. australis occurs at least as far south as southwestern Guatemala, while D. coccinea and D. imperialis also occur in parts of Central America and northern South America. Dahlia is a genus of the uplands and mountains, being found at elevations between 1,500 and 3,700 m (4,900 and 12,100 ft), in what has been described as a "pine-oak woodland" vegetative zone. Most species have limited ranges scattered throughout many mountain ranges in Mexico

Ecology
The most common pollinators are bees and small beetles.

Slugs and snails are serious pests in some parts of the world, particularly in spring when new growth is emerging through the soil. Earwigs can also disfigure the blooms and foliage. The other main pests likely to be encountered are aphids (usually on young stems and immature flower buds), red spider mite (causing foliage mottling and discolouration, worse in hot and dry conditions) and capsid bugs (resulting in contortion and holes at growing tips). Diseases affecting dahlias include powdery mildew, grey mould (Botrytis cinerea), verticillium wilt, dahlia smut (Entyloma calendulae f. dahliae), phytophthora and some plant viruses. Dahlias are a source of food for the larvae of some Lepidoptera species including angle shades, common swift, ghost moth and large yellow underwing.

Cultivation
Dahlias grow naturally in climates that do not experience frost (the tubers are hardy to USDA Zone 8), consequently they are not adapted to withstand sub-zero temperatures. However, their tuberous nature enables them to survive periods of dormancy, and this characteristic means that gardeners in temperate climates with frosts can grow dahlias successfully, provided the tubers are lifted from the ground and stored in cool yet frost-free conditions during the winter. Planting the tubers quite deep (10–15 cm or 4–6 in) also provides some protection. When in active growth, modern dahlia hybrids perform most successfully in well-watered yet free-draining soils, in situations receiving plenty of sunlight. Taller cultivars usually require some form of staking as they grow, and all garden dahlias need deadheading regularly, once flowering commences.

Horticultural classification
Horticulturally the garden dahlia is usually treated as the cultigen D. variabilis hort., which while being responsible for thousands of cultivars has an obscure taxonomic status.: 41–42 

History
The inappropriate term D. variabilis is often used to describe the cultivars of Dahlia since the correct parentage remains obscure, but probably involves Dahlia coccinea. In 1846 the Caledonia Horticultural Society of Edinburgh offered a prize of 2,000 pounds to the first person succeeding in producing a blue dahlia. This has to date not been accomplished. While dahlias produce anthocyanin, an element necessary for the production of the blue, to achieve a true blue color in a plant, the anthocyanin delphinidin needs six hydroxyl groups. To date, dahlias have only developed five, so the closest that breeders have come to achieving a "blue" specimen are variations of mauve, purples and lilac hues.

By the beginning of the twentieth century, a number of different types were recognised. These terms were based on shape or colour, and the National Dahlia Society included cactus, pompon, single, show and fancy in its 1904 guide. Many national societies developed their own classification systems until 1962 when the International Horticultural Congress agreed to develop an internationally recognised system at its Brussels meeting that year, and subsequently in Maryland in 1966. This culminated in the 1969 publication of The International Register of Dahlia Names by the Royal Horticultural Society which became the central registering authority.

This system depended primarily on the visibility of the central disc, whether it was open-centred or whether only ray florets were apparent centrally (double bloom). The double-bloom cultivars were then subdivided according to the way in which they were folded along their longitudinal axis: flat, involute (curled inwards) or revolute (curling backwards). If the end of the ray floret was split, they were considered fimbriated. Based on these characteristics, nine groups were defined plus a tenth miscellaneous group for any cultivars not fitting the above characteristics. Fimbriated dahlias were added in 2004, and two further groups (Single and Double orchid) in 2007. The last group to be added, Peony, first appeared in 2012.

In many cases the bloom diameter was then used to further label certain groups from miniature to giant. This practice was abandoned in 2012.

Modern system (RHS)
There are now more than 57,000 registered cultivars, which are officially registered through the Royal Horticultural Society (RHS). The official register is The International Register of Dahlia Names 1969 (1995 reprint) which is updated by annual supplements. The original 1969 registry published about 14,000 cultivars adding a further 1700 by 1986 and in 2003 there were 18,000. Since then about a hundred new cultivars are added annually.

Flower type
The official RHS classification lists fourteen groups, grouped by flower type, together with the abbreviations used by the RHS;

Flower size
Earlier versions of the registry subdivided some groups by flower size. Groups 4, 5, 8 and 9 were divided into five subgroups (A to E) from Giant to Miniature, and Group 6 into two subgroups, Small and Miniature. Dahlias were then described by Group and Subgroup, e.g. 5(d) ‘Ace Summer Sunset’. Some Dahlia Societies have continued this practice, but this is neither official nor standardised. As of 2013 The RHS uses two size descriptors

Dwarf Bedder (Dw.B.) – not usually exceeding 600 mm (24 in) in height, e.g. 'Preston Park' (Sin/DwB)
Lilliput dahlias (Lil) – not usually exceeding 300 mm (12 in) in height, with single, semi-double or double florets up to 26 mm (1.0 in) in diameter. ("baby" or "top-mix" dahlias), e.g. 'Harvest Tiny Tot' (Misc/Lil)
Sizes can range from tiny micro dahlias with flowers less than 50mm to giants that are over 250mm in diameter. The groupings listed here are from the New Zealand Society:

Giant-flowered cultivars have blooms with a diameter over 250 mm.
Large-flowered cultivars have blooms with a diameter of 200–250 mm.
Medium-flowered cultivars have blooms with a diameter of 155–350 mm.
Small-flowered cultivars have blooms with a diameter of 115–155 mm.
Miniature-flowered cultivars have blooms with a diameter of 50–115 mm.
Pompom-flowered cultivars have blooms with a diameter less than 50 mm.
In addition to the official classification and the terminology used by various dahlia societies, individual horticulturalists use a wide range of other descriptions, such as 'Incurved' and abbreviations in their catalogues, such as CO for Collarette.

Some plant growers include their brand name in the cultivar name. Thus Fides (part of the Dümmen Orange Group) in the Netherlands developed a series of cultivars which they named the Dahlinova series, for example Dahlinova 'Carolina Burgundy'. These are Group 10 Miscellaneous in the RHS classification scheme.

Double dahlias
In 1805, several new species were reported with red, purple, lilac, and pale yellow coloring, and the first true double flower was produced in Belgium. One of the more popular concepts of dahlia history, and the basis for many different interpretations and confusion, is that all the original discoveries were single-flowered types, which, through hybridization and selective breeding, produced double forms. Many of the species of dahlias then, and now, have single-flowered blooms. D. coccinea, the third dahlia to bloom in Europe, was a single. But two of the three drawings of dahlias by Dominguez, made in Mexico between 1570 and 1577, showed definite characteristics of doubling. In the early days of the dahlia in Europe, the word "double" simply designated flowers with more than one row of petals. The greatest effort was now directed to developing improved types of double dahlias.

During the years 1805 to 1810 several people claimed to have produced a double dahlia. In 1805 Henry C. Andrews made a drawing of such a plant in the collection of Lady Holland, grown from seedlings sent that year from Madrid. Like other doubles of the time it did not resemble the doubles of today. The first modern double, or full double, appeared in Belgium; M. Donckelaar, Director of the Botanic Garden at Louvain, selected plants for that characteristic, and within a few years secured three fully double forms. By 1826 double varieties were being grown almost exclusively, and there was very little interest in the single forms. Up to this time all the so-called double dahlias had been purple, or tinged with purple, and it was doubted if a variety untinged with that color was obtainable.

In 1843, scented single forms of dahlias were first reported in Neu Verbass, Austria. D. crocea, a fragrant variety grown from one of the Humboldt seeds, was probably interbred with the single D. coccinea. A new scented species would not be introduced until the next century when the D. coronata was brought from Mexico to Germany in 1907.

The exact date the dahlia was introduced in the United States is uncertain. One of the first dahlias in the USA may have been the D. coccinea speciosissima grown by William Leathe, of Cambridgeport, near Boston, around 1929. According to Edward Sayers, "it attracted much admiration, and at that time was considered a very elegant flower, it was however soon eclipsed by that splendid scarlet, the Countess of Liverpool". However, 9 cultivars were already listed in the catalog from Thornburn, 1825. And even earlier reference can be found in a catalogue from the Linnaean Botanical Garden, New York, 1820, that includes one scarlet, one purple, and two double orange Dahlias for sale.

Sayers stated that "No person has done more for the introduction and advancement of the culture of the Dahlia than George C. Thorburn, of New York, who yearly flowers many thousand plants at his place at Hallet's Cove, near Harlaem. The show there in the flowering season is a rich treat for the lovers of floriculture : for almost every variety can be seen growing in two large blocks or masses which lead from the road to the dwelling-house, and form a complete field of the Dahlia as a foreground to the house. Mr. T. Hogg, William Read, and many other well-known florists have also contributed much in the vicinity of New York, to the introduction of the Dahlia. Indeed so general has become the taste that almost every garden has its show of the Dahlia in the season." In Boston too there were many collections, a collection from the Messrs Hovey of Cambridgeport was also mentioned.

In 1835 Thomas Bridgeman, published a list of 160 double dahlias in his "Florist's Guide". 60 of the choicest were supplied by Mr. G. C. Thornburn of Astoria, New York, who got most of them from contacts in the UK. Not a few of them had taken prizes "at the English and American exhibitions".

"Stars of the Devil"
In 1872 J. T. van der Berg of Utrecht in the Netherlands received a shipment of seeds and plants from a friend in Mexico. The entire shipment was badly rotted and appeared to be ruined, but van der Berg examined it carefully and found a small piece of root that seemed alive. He planted and carefully tended it; it grew into a plant that he identified as a dahlia. He made cuttings from the plant during the winter of 1872–1873. This was an entirely different type of flower, with rich, red color and a high degree of doubling. In 1874 van der Berg catalogued it for sale, calling it Dahlia juarezii to honor Mexican President Benito Pablo Juarez, who had died the year before, and described it as "...equal to the beautiful color of the red poppy. Its form is very outstanding and different in every respect of all known dahlia flowers."

This plant has perhaps had a greater influence on the popularity of the modern dahlia than any other. Called "Les Etoiles du Diable" (Stars of the Devil) in France and "Cactus dahlia" elsewhere, the edges of its petals rolled backwards, rather than forward, and this new form revolutionized the dahlia world. It was thought to be a distinct mutation since no other plant that resembled it could be found in the wild. Today it is assumed that D. juarezii had, at one time, existed in Mexico and subsequently disappeared. Nurserymen in Europe crossbred this plant with dahlias discovered earlier; the results became the progenitors of all modern dahlia hybrids today.

Award of Garden Merit (RHS)
Main article: List of Award of Garden Merit dahlias
As of 2015, 124 dahlia cultivars have gained the Royal Horticultural Society's Award of Garden Merit, including:-

"Bednall beauty"
"Bishop of Llandaff"
"Clair de lune"
"David Howard"
"Ellen Huston"
"Fascination"
"Gallery Art Deco"
"Gallery Art Nouveau"
"Glorie van Heemstede"
"Honka"
"Moonfire"
"Twyning's After Eight"
Uses
The Aztecs used dahlias to treat epilepsy, and employed the long hollow stem of the D. imperialis for water pipes. Europeans attempted to introduce the tubers as a food crop, but this was unpopular.

The dahlia is considered one of the native ingredients in Oaxacan cuisine; several cultivars are still grown especially for their large, sweet potato-like tubers. Dacopa, an intense mocha-tasting extract from the roasted tubers, is used to flavor beverages throughout Central America.

Medicine
In Europe and America, prior to the discovery of insulin in 1923, diabetics—as well as consumptives—were often given a substance called Atlantic starch or diabetic sugar, derived from inulin, a naturally occurring form of fruit sugar, extracted from dahlia tubers. Inulin is still used in clinical tests for kidney functionality.

In culture
Founded in 1936, the Bloemencorso Zundert is the largest flower parade in the world entirely made by volunteers using the dahlia. The parade takes place on the first Sunday of September in Zundert, Netherlands. The floats are large artworks made of steel wire, cardboard, papier-mâché and flowers. In the Bloemencorso Zundert, mostly dahlias are used to decorate the objects and it takes thousands of them just to cover one float. Around 8 million dahlias are needed for the entire corso. Of these, around 6 million are cultivated in Zundert.

The dahlia was declared the national flower of Mexico in 1963.

Dahlia (Asteraceae) by [email protected]

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Dahlia (Asteraceae)

Dahlia is a genus of bushy, tuberous, herbaceous perennial plants native to Mexico and Central America. As a member of the Asteraceae (former name: Compositae) family of dicotyledonous plants, its relatives include the sunflower, daisy, chrysanthemum, and zinnia. There are 49 species of dahlia, with flowers in almost every hue (except blue), with hybrids commonly grown as garden plants.

Dahlias were known to the Aztecs until their Spanish conquest, after which the plants were brought to Europe. The (high in sugar) tubers of some varieties are of value to humans.

Description
Dahlias are perennial plants with tuberous roots, though they are grown as annuals in some regions with cold winters. While some have herbaceous stems, others have stems which lignify in the absence of secondary tissue and resprout following winter dormancy, allowing further seasons of growth. As a member of the Asteraceae, the dahlia has a flower head that is actually a composite (hence the older name Compositae) with both central disc florets and surrounding ray florets. Each floret is a flower in its own right, but is often incorrectly described as a petal. The modern name Asteraceae refers to the appearance of a star with surrounding rays.

The stems are leafy, ranging in height from as low as 30 centimetres (12 inches) to more than 1.8–2.4 metres (6–8 feet). Flower forms are variable, with one head per stem; these can be as small as 5 cm (2 in) in diameter or up to 30 cm (1 ft) ("dinner plate"). The majority of species do not produce scented flowers. Like most plants that do not attract pollinating insects through scent, they are brightly colored, displaying most hues, with the exception of blue.

The great variety in species results from garden dahlias being octoploids, having eight sets of homologous chromosomes. In addition, dahlias also contain many transposons—genetic pieces that move from place to place upon an allele—which contributes to their manifesting such great diversity.

Taxonomy
Spaniards reported finding the plants growing in Mexico in 1525, but the earliest known description is by Francisco Hernández, physician to Philip II, who was ordered to visit Mexico in 1570 to study the "natural products of that country". They were used as a source of food by the indigenous peoples, who both gathered wild specimens and cultivated crops. The indigenous peoples variously identified the plants as "Chichipatl" (Toltecs) and "Acocotle" or "Cocoxochitl" (Aztecs). From Hernandez's perception of Nahuatl to Spanish (through various other translations) the word is "water cane", "water pipe", "water pipe flower", "hollow stem flower", or "cane flower", all referring to the hollow plant stems.

Hernandez described two varieties of dahlias (the pinwheel-like Dahlia pinnata and the huge Dahlia imperialis) as well as other medicinal plants of New Spain. Francisco Dominguez, an Hidalgo gentleman who accompanied Hernandez on part of his seven-year study, made a series of drawings to supplement the four volume report. Three of his drawings showed plants with flowers: two resembled the modern bedding dahlia, and one resembled the species Dahlia merckii; all displayed a high degree of doubleness. In 1578, a manuscript titled Nova Plantarum, Animalium et Mineralium Mexicanorum Historia, was sent back to the Escorial in Madrid. It was translated into Latin by Francisco Ximenes in 1615. In 1640, Francisco Cesi, President of the Academia dei Lincei of Rome, bought the Ximenes translation and, after annotating it, published it in 1649–1651 as two volumes, Rerum Medicarum Novae Hispaniae Thesaurus Seu Nova Plantarium, Animalium et Mineralium Mexicanorum Historia. The original manuscripts were destroyed in a fire in the mid-1600s.

European introduction
In 1787, the French botanist Nicolas-Joseph Thiéry de Menonville, sent to Mexico to steal the cochineal insect valued for its scarlet dye, reported the strangely beautiful flowers he had seen growing in a garden in Oaxaca. In 1789, Vicente Cervantes, director of the Botanical Garden at Mexico City, sent "plant parts" to Abbe Antonio José Cavanilles, director of the Royal Gardens of Madrid. Cavanilles flowered one plant that same year, then the second one a year later. In 1791 he called the new growths "Dahlia" for Anders Dahl.[1] The first plant was called Dahlia pinnata after its pinnate foliage; the second, Dahlia rosea for its rose-purple color. In 1796, from the parts sent by Cervantes, Cavanilles flowered a third plant, which he named Dahlia coccinea for its scarlet color.

In 1798, Cavanilles sent D. pinnata seeds to Parma, Italy. That year, the Marchioness of Bute, wife of the Earl of Bute, the English Ambassador to Spain, obtained a few seeds from Cavanilles and sent them to Kew Gardens, where they flowered but were lost after two to three years.


The Dahlia Garden at Holland House in 1907
In the following years Madrid sent seeds to Berlin and Dresden in Germany, and to Turin and Thiene in Italy. In 1802, Cavanilles sent tubers of "these three" (D. pinnata, D. rosea, D. coccinea) to Swiss botanist Augustin Pyramus de Candolle at University of Montpelier in France, Andre Thouin at the Jardin des Plantes in Paris and Scottish botanist William Aiton at Kew Gardens. That same year, John Fraser, English nurseryman and later botanical collector to the Czar of Russia, brought D. coccinea seeds from Paris to the Apothecaries Gardens in England, where they flowered in his greenhouse a year later, providing Botanical Magazine with an illustration.

In 1804, a new species, Dahlia sambucifolia, was successfully grown at Holland House, Kensington. Whilst in Madrid in 1804, Lady Holland was given either dahlia seeds or tubers by Cavanilles. She sent them back to England, to Lord Holland's librarian at Holland House, who successfully raised the plants and produced two double flowers a year later. The plants raised in 1804 did not survive; new stock was brought from France in 1815. In 1824, Lord Holland sent his wife a note containing the following verse:

The dahlia you brought to our isle
Your praises for ever shall speak;
Mid gardens as sweet as your smile,
And in colour as bright as your cheek.

In 1805, German naturalist Alexander von Humboldt sent more seeds from Mexico to Aiton in England, Thouin in Paris, and Christoph Friedrich Otto, director of the Berlin Botanical Garden. More significantly, he sent seeds to botanist Carl Ludwig Willdenow in Germany. Willdenow now reclassified the rapidly growing number of species, changing the genus from Dahlia to Georgina; after naturalist Johann Gottlieb Georgi. He combined the Cavanilles species D. pinnata and D. rosea under the name of Georgina variabilis; D. coccinea was still held to be a separate species, which he renamed Georgina coccinea.

Classification
Since 1789 when Cavanilles first flowered the dahlia in Europe, there has been an ongoing effort by many growers, botanists and taxonomists, to determine the development of the dahlia to modern times. At least 85 species have been reported: approximately 25 of these were first reported from the wild; the remainder appeared in gardens in Europe. They were considered hybrids, the results of crossing between previously reported species, or developed from the seeds sent by Humboldt from Mexico in 1805, or perhaps from some other undocumented seeds that had found their way to Europe. Several of these were soon discovered to be identical with earlier reported species, but the greatest number are new varieties. Morphological variation is highly pronounced in the dahlia. William John Cooper Lawrence, who hybridized hundreds of families of dahlias in the 1920s, stated: "I have not yet seen any two plants in the families I have raised which were not to be distinguished one from the other. Constant reclassification of the 85 reported species has resulted in a considerably smaller number of distinct species, as there is a great deal of disagreement today between systematists over classification.

In 1829, all species growing in Europe were reclassified under an all-encompassing name of D. variabilis, Desf., though this is not an accepted name. Through the interspecies cross of the Humboldt seeds and the Cavanilles species, 22 new species were reported by that year, all of which had been classified in different ways by several different taxonomists, creating considerable confusion as to which species was which. As of now Dahlias are classified into 15 different species by botanist Liberty Hyde Bailey.

In 1830 William Smith suggested that all dahlia species could be divided into two groups for color, red-tinged and purple-tinged. In investigating this idea Lawrence determined that with the exception of D. variabilis, all dahlia species may be assigned to one of two groups for flower-colour: Group I (ivory-magenta) or Group II (yellow-orange-scarlet).

Modern classification
The genus Dahlia is situated in the Asteroideae subfamily of the Asteraceae, in the Coreopsideae tribe. Within that tribe it is the second largest genus, after Coreopsis, and appears as a well defined clade within the Coreopsideae.

Subdivision
Infrageneric subdivision
See also: List of Dahlia species
Sherff (1955), in the first modern taxonomy described three sections for the 18 species he recognised, Pseudodendron, Epiphytum and Dahlia. By 1969 Sørensen recognised 29 species and four sections by splitting off Entemophyllon from section Dahlia. By contrast Giannasi (1975) using a phytochemical analysis based on flavonoids, reduced the genus to just two sections, Entemophyllon and Dahlia, the latter having three subsections, Pseudodendron, Dahlia, and Merckii. Sørensen then issued a further revision in 1980, incorporating subsection Merckii in his original section Dahlia. When he described two new species in the 1980s (Dahlia tubulata and D. congestifolia), he placed them within his existing sections. A further species, Dahlia sorensenii was added by Hansen and Hjerting in (1996). At the same time they demonstrated that Dahlia pinnata should more properly be designated D. x pinnata. D. x pinnata was shown to actually be a variant of D. sorensenii that had acquired hybrid qualities before it was introduced to Europe in the sixteenth century and formally named by Cavanilles. The original wild D. pinnata is presumed extinct. Further species continue to be described, Saar (2003) describing 35 species. However separation of the sections on morphological, cytologal and biocemical criteria has not been entirely satisfactory.

To date these sectional divisions have not been fully supported phylogenetically, which demonstrate only section Entemophyllon as a distinct sectional clade. The other major grouping is the core Dahlia clade (CDC), which includes most of the section Dahlia. The remainder of the species occupy what has been described as the variable root clade (VRC) which includes the small section Pseudodendron but also the monotypic section Epiphytum and a number of species from within section Dahlia. Outside of these three clades lie D. tubulata and D. merckii as a polytomy.

Horticulturally the sections retain some usage, section Pseudodendron being referred to as 'Tree Dahlias', Epiphytum as the 'Vine Dahlia'. The remaining two herbaceous sections being distinguished by their pinnules, opposing (Dahlia) or alternating (Entemophyllon).

Sections
Sections (including chromosome numbers), with geographical distribution;

Epiphytum Sherff (2n = 32)
10 m tall climber with aerial roots 5 cm thick and up to more than 20 m long; pinnules opposite
1 species, D. macdougallii Sherff
Mexico: Oaxaca
Entemophyllon P. D. Sorensen (2n = 34)
6 species
Mexico: Hidalgo, Nuevo León, Tamaulipas, Querétaro, Durango, San Luis Potosí
Pseudodendron P. D. Sorensen (2n = 32)
3 species + D. excelsa of uncertain identity
Mexico: Chiapas, Guerrero, Jalisco, Michoacan, Oaxaca, and Costa Rica, El Salvador, Guatemala & Colombia
Dahlia (2n = 32, 36 or 64)
24 species
Mexico: Distrito Federal, Guerrero, Hidalgo, Morelos, Nuevo León, Puebla, San Luis Potosí, Tamaulipas, Veracruz, Oaxaca, Puebla, Chiapas, México, Huehuetenango, Chihuahua, Durango, Michoacan & Guatemala
Only Pseudodendron (D. imperialis) and Dahlia (D. australis, D. coccinea) occur outside Mexico.

Species
Main article: List of Dahlia species
There are currently 42 accepted species in the genus Dahlia but new species continue to be described.

Etymology
The naming of the plant itself has long been a subject of some confusion. Many sources state that the name "Dahlia" was bestowed by the pioneering Swedish botanist and taxonomist Carl Linnaeus to honor his late student, Anders Dahl, author of Observationes Botanicae. However, Linnaeus died in 1778, more than eleven years before the plant was introduced into Europe in 1789, so while it is generally agreed that the plant was named in 1791 in honor of Dahl, who had died two years before, Linnaeus could not have been the one who did so. It was probably Abbe Antonio Jose Cavanilles, Director of the Royal Gardens of Madrid, who should be credited with the attempt to scientifically define the genus, since he not only received the first specimens from Mexico in 1789, but named the first three species that flowered from the cuttings.

Regardless of who bestowed it, the name was not so easily established. In 1805, German botanist Carl Ludwig Willdenow, asserting that the genus Dahlia Thunb. (published a year after Cavanilles's genus and now considered a synonym of Trichocladus) was more widely accepted, changed the plants' genus from Dahlia to Georgina (after the German-born naturalist Johann Gottlieb Georgi, a professor at the Imperial Academy of Sciences of St. Petersburg, Russia). He also reclassified and renamed the first three species grown, and identified, by Cavanilles. It was not until 1810, in a published article, that he officially adopted the Cavanilles's original designation of Dahlia. However, the name Georgina still persisted in Germany for the next few decades. In Russian, it is still named Georgina (Russian: георгинa).

Distribution and habitat
Dahlia is found predominantly in Mexico, but some species are found ranging as far south as northern South America. D. australis occurs at least as far south as southwestern Guatemala, while D. coccinea and D. imperialis also occur in parts of Central America and northern South America. Dahlia is a genus of the uplands and mountains, being found at elevations between 1,500 and 3,700 m (4,900 and 12,100 ft), in what has been described as a "pine-oak woodland" vegetative zone. Most species have limited ranges scattered throughout many mountain ranges in Mexico

Ecology
The most common pollinators are bees and small beetles.

Pests and diseases
Main article: List of Dahlia diseases
Slugs and snails are serious pests in some parts of the world, particularly in spring when new growth is emerging through the soil. Earwigs can also disfigure the blooms and foliage. The other main pests likely to be encountered are aphids (usually on young stems and immature flower buds), red spider mite (causing foliage mottling and discolouration, worse in hot and dry conditions) and capsid bugs (resulting in contortion and holes at growing tips). Diseases affecting dahlias include powdery mildew, grey mould (Botrytis cinerea), verticillium wilt, dahlia smut (Entyloma calendulae f. dahliae), phytophthora and some plant viruses. Dahlias are a source of food for the larvae of some Lepidoptera species including angle shades, common swift, ghost moth and large yellow underwing.

Cultivation
Dahlias grow naturally in climates that do not experience frost (the tubers are hardy to USDA Zone 8), consequently they are not adapted to withstand sub-zero temperatures. However, their tuberous nature enables them to survive periods of dormancy, and this characteristic means that gardeners in temperate climates with frosts can grow dahlias successfully, provided the tubers are lifted from the ground and stored in cool yet frost-free conditions during the winter. Planting the tubers quite deep (10–15 cm or 4–6 in) also provides some protection. When in active growth, modern dahlia hybrids perform most successfully in well-watered yet free-draining soils, in situations receiving plenty of sunlight. Taller cultivars usually require some form of staking as they grow, and all garden dahlias need deadheading regularly, once flowering commences.

History
The inappropriate term D. variabilis is often used to describe the cultivars of Dahlia since the correct parentage remains obscure, but probably involves Dahlia coccinea. In 1846 the Caledonia Horticultural Society of Edinburgh offered a prize of 2,000 pounds to the first person succeeding in producing a blue dahlia. This has to date not been accomplished. While dahlias produce anthocyanin, an element necessary for the production of the blue, to achieve a true blue color in a plant, the anthocyanin delphinidin needs six hydroxyl groups. To date, dahlias have only developed five, so the closest that breeders have come to achieving a "blue" specimen are variations of mauve, purples and lilac hues.

By the beginning of the twentieth century, a number of different types were recognised. These terms were based on shape or colour, and the National Dahlia Society included cactus, pompon, single, show and fancy in its 1904 guide. Many national societies developed their own classification systems until 1962 when the International Horticultural Congress agreed to develop an internationally recognised system at its Brussels meeting that year, and subsequently in Maryland in 1966. This culminated in the 1969 publication of The International Register of Dahlia Names by the Royal Horticultural Society which became the central registering authority.

This system depended primarily on the visibility of the central disc, whether it was open-centred or whether only ray florets were apparent centrally (double bloom). The double-bloom cultivars were then subdivided according to the way in which they were folded along their longitudinal axis: flat, involute (curled inwards) or revolute (curling backwards). If the end of the ray floret was split, they were considered fimbriated. Based on these characteristics, nine groups were defined plus a tenth miscellaneous group for any cultivars not fitting the above characteristics. Fimbriated dahlias were added in 2004, and two further groups (Single and Double orchid) in 2007. The last group to be added, Peony, first appeared in 2012.

In many cases the bloom diameter was then used to further label certain groups from miniature to giant. This practice was abandoned in 2012.

Modern system (RHS)
There are now more than 57,000 registered cultivars, which are officially registered through the Royal Horticultural Society (RHS). The official register is The International Register of Dahlia Names 1969 (1995 reprint) which is updated by annual supplements. The original 1969 registry published about 14,000 cultivars adding a further 1700 by 1986 and in 2003 there were 18,000. Since then about a hundred new cultivars are added annually.[

Flower type
The official RHS classification lists fourteen groups, grouped by flower type, together with the abbreviations used by the RHS;

Group 1 – Single-flowered dahlias (Sin) – Flower has a central disc with a single outer ring of florets (which may overlap) encircling it, and which may be rounded or pointed.
(e.g. 'Twyning's After Eight')

Group 2 – Anemone-flowered dahlias (Anem) – The centre of the flower consists of dense elongated tubular florets, longer than the disc florets of Single dahlias, while the outer parts have one or more rings of flatter ray florets. Disc absent.
(e.g. 'Boogie Woogie')
'
Group 3 – Collerette dahlias (Col) – Large flat florets forming a single outer ring around a central disc and which may overlap a smaller circle of florets closer to the centre, which have the appearance of a collar.
(e.g. 'Starsister', 'Lilian Alice', 'Apple Blossom')
'
Group 4 – Waterlily dahlias (WL) – Double blooms, broad sparse curved, slightly curved or flat florets and very shallow in depth compared with other dahlias. Depth less than half the diameter of the bloom.
(e.g. 'Cameo')

Group 5 – Decorative dahlias (D) – Double blooms, ray florets broad, flat, involute no more than seventy five per cent of the longitudinal axis, slightly twisted and usually bluntly pointed. No visible central disc.
(e.g. 'Berliner Kleene')

Group 6 – Ball dahlias (Ba) – Double blooms that are ball shaped or slightly flattened. Ray florets blunt or rounded at the tips, margins arranged spirally, involute for at least seventy five percent of the length of the florets. Larger than Pompons.
(e.g. 'Barbarry Ball')

Group 7 – Pompon dahlias (Pom) – Double spherical miniature flowers made up entirely from florets that are curved inwards (involute) for their entire length (longitudinal axis), resembling a pompon.
(e.g. 'Small World')

Group 8 – Cactus dahlias (C) – Double blooms, ray florets pointed, with majority revolute (rolled) over more than fifty percent of their longitudinal axis, and straight or incurved. Narrower than Semi cactus.
(e.g. 'Nuit d'Eté')

Group 9 – Semi cactus dahlias (S–c) – Double blooms, very pointed ray florets, revolute for greater than twenty five percent and less than fifty percent of their longitudinal axis. Broad at the base and straight or incurved, almost spiky in appearance.
(e.g. 'Mick's Peppermint')
'Mick's Peppermint' (Semi Cactus)
'
Group 10 – Miscellaneous dahlias (Misc) – not described in any other group.
(e.g. 'Moonfire', 'Carolina Burgundy')
'

Hybrid
Group 11 – Fimbriated dahlias (Fim) – ray florets evenly split or notched into two or more divisions, uniformly throughout the bloom, creating a fimbriated (fringed) effect. The petals may be flat, involute, revolute, straight, incurving or twisted.
(e.g. 'Marlene Joy')
'Marlene Joy' (Fimbriated)

Group 12 – Single Orchid (Star) dahlias (SinO) – single outer ring of florets surround a central disc. The ray florets are either involute or revolute.
(e.g. 'Alloway Candy)
'Alloway Candy' (Single Orchid (Star))

Group 13 – Double Orchid dahlias (DblO) – Double blooms with triangular centres. The ray florets are narrowly lanceolate and are either involute or revolute. The central disc is absent.
(e.g. 'Pink Giraffe')
'Pink Giraffe' (Double Orchid)

Group 14 – Peony-flowered dahlias (P) – Large flowers with three or four rows of rays that are flattened and expanded and arranged irregularly. The rays surround a golden disc similar to that of Single dahlias.
(e.g. 'Bishop of Llandaff')

Flower size
Earlier versions of the registry subdivided some groups by flower size. Groups 4, 5, 8 and 9 were divided into five subgroups (A to E) from Giant to Miniature, and Group 6 into two subgroups, Small and Miniature. Dahlias were then described by Group and Subgroup, e.g. 5 ‘Ace Summer Sunset’. Some Dahlia Societies have continued this practice, but this is neither official nor standardised. As of 2013 The RHS uses two size descriptors

Dwarf Bedder (Dw.B.) – not usually exceeding 600 mm (24 in) in height, e.g. 'Preston Park' (Sin/DwB)
Lilliput dahlias (Lil) – not usually exceeding 300 mm (12 in) in height, with single, semi-double or double florets up to 26 mm (1.0 in) in diameter. ("baby" or "top-mix" dahlias), e.g. 'Harvest Tiny Tot' (Misc/Lil)
Sizes can range from tiny micro dahlias with flowers less than 50mm to giants that are over 250mm in diameter. The groupings listed here are from the New Zealand Society:

Giant-flowered cultivars have blooms with a diameter over 250 mm.
Large-flowered cultivars have blooms with a diameter of 200–250 mm.
Medium-flowered cultivars have blooms with a diameter of 155–350 mm.
Small-flowered cultivars have blooms with a diameter of 115–155 mm.
Miniature-flowered cultivars have blooms with a diameter of 50–115 mm.
Pompom-flowered cultivars have blooms with a diameter less than 50 mm.
In addition to the official classification and the terminology used by various dahlia societies, individual horticulturalists use a wide range of other descriptions, such as 'Incurved' and abbreviations in their catalogues, such as CO for Collarette.

Branding
Some plant growers include their brand name in the cultivar name. Thus Fides (part of the Dümmen Orange Group) in the Netherlands developed a series of cultivars which they named the Dahlinova series, for example Dahlinova 'Carolina Burgundy'. These are Group 10 Miscellaneous in the RHS classification scheme.

Double dahlias
In 1805, several new species were reported with red, purple, lilac, and pale yellow coloring, and the first true double flower was produced in Belgium. One of the more popular concepts of dahlia history, and the basis for many different interpretations and confusion, is that all the original discoveries were single-flowered types, which, through hybridization and selective breeding, produced double forms Many of the species of dahlias then, and now, have single-flowered blooms. D. coccinea, the third dahlia to bloom in Europe, was a single. But two of the three drawings of dahlias by Dominguez, made in Mexico between 1570 and 1577, showed definite characteristics of doubling. In the early days of the dahlia in Europe, the word "double" simply designated flowers with more than one row of petals. The greatest effort was now directed to developing improved types of double dahlias.

During the years 1805 to 1810 several people claimed to have produced a double dahlia. In 1805 Henry C. Andrews made a drawing of such a plant in the collection of Lady Holland, grown from seedlings sent that year from Madrid. Like other doubles of the time it did not resemble the doubles of today. The first modern double, or full double, appeared in Belgium; M. Donckelaar, Director of the Botanic Garden at Louvain, selected plants for that characteristic, and within a few years secured three fully double forms. By 1826 double varieties were being grown almost exclusively, and there was very little interest in the single forms. Up to this time all the so-called double dahlias had been purple, or tinged with purple, and it was doubted if a variety untinged with that color was obtainable.

In 1843, scented single forms of dahlias were first reported in Neu Verbass, Austria. D. crocea, a fragrant variety grown from one of the Humboldt seeds, was probably interbred with the single D. coccinea. A new scented species would not be introduced until the next century when the D. coronata was brought from Mexico to Germany in 1907.

The exact date the dahlia was introduced in the United States is uncertain. One of the first dahlias in the USA may have been the D. coccinea speciosissima grown by William Leathe, of Cambridgeport, near Boston, around 1929. According to Edward Sayers, "it attracted much admiration, and at that time was considered a very elegant flower, it was however soon eclipsed by that splendid scarlet, the Countess of Liverpool". However, 9 cultivars were already listed in the catalog from Thornburn, 1825. And even earlier reference can be found in a catalogue from the Linnaean Botanical Garden, New York, 1820, that includes one scarlet, one purple, and two double orange Dahlias for sale.

Sayers stated that "No person has done more for the introduction and advancement of the culture of the Dahlia than George C. Thorburn, of New York, who yearly flowers many thousand plants at his place at Hallet's Cove, near Harlaem. The show there in the flowering season is a rich treat for the lovers of floriculture : for almost every variety can be seen growing in two large blocks or masses which lead from the road to the dwelling-house, and form a complete field of the Dahlia as a foreground to the house. Mr. T. Hogg, William Read, and many other well-known florists have also contributed much in the vicinity of New York, to the introduction of the Dahlia. Indeed so general has become the taste that almost every garden has its show of the Dahlia in the season." In Boston too there were many collections, a collection from the Messrs Hovey of Cambridgeport was also mentioned.

In 1835 Thomas Bridgeman, published a list of 160 double dahlias in his "Florist's Guide". 60 of the choicest were supplied by Mr. G. C. Thornburn of Astoria, New York, who got most of them from contacts in the UK. Not a few of them had taken prizes "at the English and American exhibitions".

"Stars of the Devil"
In 1872 J. T. van der Berg of Utrecht in the Netherlands received a shipment of seeds and plants from a friend in Mexico. The entire shipment was badly rotted and appeared to be ruined, but van der Berg examined it carefully and found a small piece of root that seemed alive. He planted and carefully tended it; it grew into a plant that he identified as a dahlia. He made cuttings from the plant during the winter of 1872–1873. This was an entirely different type of flower, with rich, red color and a high degree of doubling. In 1874 van der Berg catalogued it for sale, calling it Dahlia juarezii to honor Mexican President Benito Pablo Juarez, who had died the year before, and described it as "...equal to the beautiful color of the red poppy. Its form is very outstanding and different in every respect of all known dahlia flowers."

This plant has perhaps had a greater influence on the popularity of the modern dahlia than any other. Called "Les Etoiles du Diable" (Stars of the Devil) in France and "Cactus dahlia" elsewhere, the edges of its petals rolled backwards, rather than forward, and this new form revolutionized the dahlia world. It was thought to be a distinct mutation since no other plant that resembled it could be found in the wild. Today it is assumed that D. juarezii had, at one time, existed in Mexico and subsequently disappeared. Nurserymen in Europe crossbred this plant with dahlias discovered earlier; the results became the progenitors of all modern dahlia hybrids today.

Award of Garden Merit (RHS)
Main article: List of Award of Garden Merit dahlias
As of 2015, 124 dahlia cultivars have gained the Royal Horticultural Society's Award of Garden Merit, including:-

"Bednall beauty"
"Bishop of Llandaff"
"Clair de lune"
"David Howard"
"Ellen Huston"
"Fascination"
"Gallery Art Deco"
"Gallery Art Nouveau"
"Glorie van Heemstede"
"Honka"
"Moonfire"
"Twyning's After Eight"

Uses
The Aztecs used dahlias to treat epilepsy, and employed the long hollow stem of the D. imperialis for water pipes. Europeans attempted to introduce the tubers as a food crop, but this was unpopular.

The dahlia is considered one of the native ingredients in Oaxacan cuisine; several cultivars are still grown especially for their large, sweet potato-like tubers. Dacopa, an intense mocha-tasting extract from the roasted tubers, is used to flavor beverages throughout Central America.

Medicine
In Europe and America, prior to the discovery of insulin in 1923, diabetics—as well as consumptives—were often given a substance called Atlantic starch or diabetic sugar, derived from inulin, a naturally occurring form of fruit sugar, extracted from dahlia tubers. Inulin is still used in clinical tests for kidney functionality.

In culture
Founded in 1936, the Bloemencorso Zundert is the largest flower parade in the world entirely made by volunteers using the dahlia. The parade takes place on the first Sunday of September in Zundert, Netherlands. The floats are large artworks made of steel wire, cardboard, papier-mâché and flowers. In the Bloemencorso Zundert, mostly dahlias are used to decorate the objects and it takes thousands of them just to cover one float. Around 8 million dahlias are needed for the entire corso. Of these, around 6 million are cultivated in Zundert.

The dahlia was declared the national flower of Mexico in 1963.

Tagete (Asteraceae) by [email protected]

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Tagete (Asteraceae)

Tagetes is a genus of 50 species of annual or perennial, mostly herbaceous plants in the family Asteraceae. They are among several groups of plants known in English as marigolds. The genus Tagetes was described by Carl Linnaeus in 1753.

These plants are native to Mexico, growing naturally from Mexico's valley down to the south and even reaching several other Latinamerican countries, but some species have become naturalized around the world. One species, T. minuta, is considered a noxious invasive plant in some areas.

Description
Tagetes species vary in size from 0.1 to 2.2 m tall. Most species have pinnate green leaves. Blooms naturally occur in golden, orange, yellow, and white colors, often with maroon highlights. Floral heads are typically (1-) to 4–6 cm diameter, generally with both ray florets and disc florets. In horticulture, they tend to be planted as annuals, although the perennial species are gaining popularity. They have fibrous roots.

Depending on the species, Tagetes species grow well in almost any sort of soil. Most horticultural selections grow best in soil with good drainage, and some cultivars are known to have good tolerance to drought.

Nomenclature
The Latin Tagētes derives from the Tages in Etruscan mythology, born from plowing the earth. It likely refers to the ease with which plants of this genus come out each year either by the seeds produced in the previous year, or by the stems which regrow from the stump already in place.

The common name in English, marigold, is derived from Mary's gold, a name first applied to a similar plant native to Europe, Calendula officinalis.

The most commonly cultivated varieties of Tagetes are known variously as African marigolds (usually referring to cultivars and hybrids of Tagetes erecta), or French marigolds (usually referring to hybrids and cultivars of Tagetes patula, many of which were developed in France). The so-called signet marigolds are hybrids derived mostly from Tagetes tenuifolia.

Cultivation and uses
Depending on the species, marigold foliage has a musky, pungent scent, though some varieties have been bred to be scentless. Due to antibacterial thiophenes exuded by the roots, Tagetes should not be planted near any legume crop.[11] Some of the perennial species are deer-, rabbit-, rodent- and javelina or peccary-resistant.

T. minuta (khakibush or huacatay), originally from South America, has been used as a source of essential oil for the perfume and industry known as tagette or "marigold oil", and as a flavourant in the food and tobacco industries. It is commonly cultivated in South Africa, where the species is also a useful pioneer plant in the reclamation of disturbed land.

The florets of Tagetes erecta are rich in the orange-yellow carotenoid lutein and are used as a food colour (INS number E161b) in the European Union for foods such as pasta, vegetable oil, margarine, mayonnaise, salad dressing, baked goods, confectionery, dairy products, ice cream, yogurt, citrus juice and mustard. In the United States, however, the powders and extracts are only approved as colorants in poultry feed.

Marigolds are recorded as a food plant for some Lepidoptera caterpillars including the dot moth, and a nectar source for other butterflies and bumblebees. They are often part of butterfly gardening plantings. In the wild, many species are pollinated by beetles.

Cultural significance
Tagetes lucida
The species Tagetes lucida, known as pericón, is used to prepare a sweetish, anise-flavored medicinal tea in Mexico. It is also used as a culinary herb in many warm climates, as a substitute for tarragon, and offered in the nursery as "Texas tarragon" or "Mexican mint marigold".

Tagetes minuta, native to southern South America, is a tall, upright marigold plant with small flowers used as a culinary herb in Peru, Ecuador, and parts of Chile and Bolivia, where it is called by the Incan term huacatay. The paste is used to make the popular potato dish called ocopa. Having both "green" and "yellow/orange" notes, the taste and odor of fresh T. minuta is like a mixture of sweet basil, tarragon, mint and citrus. It is also used as a medicinal tea for gastrointestinal complaints and specifically against nematodes.

Tagetes erecta
Tagetes erecta is widely used in Day of the Dead celebrations in Mexico.

Tagetes – various species
In Bangladesh, India and other South Asian countries, marigold is used for ornamentation purposes in functions like turmeric ceremony, weddings, Pohela Falgun and other functions. During the colonial period the native varieties of these flowers were replaced by American species like T. erecta, T. patula and T. tenuifolia. The marigold is also widely cultivated in India and Thailand, particularly the species T. erecta, Tagetes patula and T. tenuifolia. It is always sold in the markets for daily worship and rituals. Vast quantities of marigolds are used in garlands and decoration for weddings, festivals, and religious events. Marigold cultivation is extensively seen in Telangana, Andhra Pradesh, Tamil Nadu, West Bengal, Karnataka and Uttar Pradesh (for the Vijayadashami and Diwali markets) states of India.

In Ukraine, chornobryvtsi (T. erecta, T. patula and the signet marigold, l. tenuifolia) are regarded as one of the national symbols, and are often mentioned in songs, poems and tales.[citation needed]


Species
Accepted species
Tagetes apetala
Tagetes arenicola
Tagetes argentina Cabrera
Tagetes biflora Cabrera
Tagetes campanulata Griseb.
Tagetes daucoides
Tagetes elliptica Sm.
Tagetes elongata
Tagetes epapposa
Tagetes erecta L. – African marigold, Aztec marigold
Tagetes filifolia Lag.
Tagetes foeniculacea
Tagetes foetidissima DC.
Tagetes hartwegii Greenm.
Tagetes iltisiana Rydb.
Tagetes inclusa
Tagetes lacera
Tagetes laxa Cabrera
Tagetes lemmonii A.Gray – Mt. Lemmon marigold, Mexican marigold
Tagetes linifolia
Tagetes lucida Cav. – Mexican mint marigold, Texas tarragon
Tagetes mandonii
Tagetes mendocina Phil.
Tagetes micrantha Cav. – licorice marigold
Tagetes microglossa
Tagetes minima
Tagetes minuta L. – wild marigold
Tagetes moorei
Tagetes mulleri S.F.Blake
Tagetes multiflora Kunth
Tagetes nelsonii Greenm.
Tagetes oaxacana
Tagetes osteni
Tagetes palmeri
Tagetes parryi A.Gray
Tagetes patula L. – French marigold
Tagetes perezii Cabrera
Tagetes praetermissa
Tagetes pringlei S.Watson
Tagetes pusilla
Tagetes riojana M.Ferraro
Tagetes rupestris Cabrera
Tagetes stenophylla B.L.Rob.
Tagetes subulata Cerv.
Tagetes subvillosa
Tagetes tenuifolia Cav. – signet marigold
Tagetes terniflora Kunth
Tagetes triradiata
Tagetes verticillata Lag. & Rodr.
Tagetes zypaquirensis Humb. & Bonpl.

Bellis (Asteraceae) by [email protected]

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Bellis (Asteraceae)

Bellis is a genus of flowering plants in the family Asteraceae.

The group is native to Europe, the Mediterranean and northern Africa. One species has been introduced into North America and others into other parts of the world. The genus includes the familiar common daisy Bellis perennis.

Description
Bellis species are mostly perennials, and grow from 5–20 cm (2–8 in) tall. They have simple erect stems, and most species have basal leaves. They have radiate flower heads that are produced one per stem.

Species
Bellis annua L.
Bellis annua subsp. microcephala
Bellis annua subsp. minuta
Bellis azorica L.
Bellis bernardii Boiss. & Reut.
Bellis caerulescens (Coss.) Coss. ex Ball
Bellis cordifolia (Kunze) Willk.
Bellis dubia Spreng.
Bellis hyrcanica Woronow
Bellis longifolia Boiss. & Heldr. (synonym of Bellis sylvestris)
Bellis microcephala Lange (synonym of Bellis annua subsp. microcephala)
Bellis pappulosa Boiss. ex DC.
Bellis perennis L.
Bellis rotundifolia (Desf.) Boiss. & Reut.
Bellis sylvestris (L.) Cyr.
Bellis sylvestris subsp. pappulosa

Possible infrageneric groups
Bellis sect. Brachyscome (Cass.) Baill.
Bellis sect. Brachystephium (Less.) Kuntze
Bellis sect. Lagenophora (Cass.) Baill.
Bellis sect. Silphiosperma (Steetz) Kuntze

Common Daisy - Bellis Perennis (Bellis) by [email protected]

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Common Daisy - Bellis Perennis (Bellis)

Bellis is a genus of flowering plants in the family Asteraceae.

The group is native to Europe, the Mediterranean and northern Africa. One species has been introduced into North America and others into other parts of the world. The genus includes the familiar common daisy Bellis perennis.

Description
Bellis species are mostly perennials, and grow from 5–20 cm (2–8 in) tall. They have simple erect stems, and most species have basal leaves. They have radiate flower heads that are produced one per stem.

Species
Accepted species
Bellis annua L.
Bellis annua subsp. microcephala
Bellis annua subsp. minuta
Bellis azorica L.
Bellis bernardii Boiss. & Reut.
Bellis caerulescens (Coss.) Coss. ex Ball
Bellis cordifolia (Kunze) Willk.
Bellis dubia Spreng.
Bellis hyrcanica Woronow
Bellis longifolia Boiss. & Heldr. (synonym of Bellis sylvestris)
Bellis microcephala Lange (synonym of Bellis annua subsp. microcephala)
Bellis pappulosa Boiss. ex DC.
Bellis perennis L.
Bellis rotundifolia (Desf.) Boiss. & Reut.
Bellis sylvestris (L.) Cyr.
Bellis sylvestris subsp. pappulosa

Dahlia - Garden Dahlia (Dahlia Pinnata) by [email protected]

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Dahlia - Garden Dahlia (Dahlia Pinnata)

Dahlia pinnata is a species in the genus Dahlia, family Asteraceae, with the common name garden dahlia. It is the type species of the genus and is widely cultivated.

Description
Dahlia pinnata is a perennial herbaceous plant with a rhizome and tuberous roots, reaching a height of 70 to 120 centimetres (28 to 47 inches), rarely 160 cm. The stem is erect, and branched only in the inflorescence. The leaves are usually simple, with leaflets that are ovate and 5–10 centimetres (2–4 in) long. The plant is slightly shaggy.

Between July and October, atop stems 5 to 15 cm in length are two to eight flower heads, 6 to 10 cm in diameter. The eight florets have a length of 3 to 5 cm, are ovate and coloured pink to deep purple.

Taxonomy
Hansen and Hjerting in (1996) demonstrated that Dahlia pinnata should more properly be designated D. x pinnata. D. x pinnata was shown to actually be a variant of D. sorensenii that had acquired hybrid qualities before it was introduced to Europe in the sixteenth century and formally named by Cavanilles. The original wild D. pinnata is presumed extinct.

Distribution and habitat
It is geographically located in Central America, tending to grow at borders. The plant occurs in Mexico in the mountains around Mexico City.

Ecology
Dahlias tend to attract quite a bit of insects, some which are dangerous and harmful to their survival. Insects like slugs, earwigs, the red spider, snails, caterpillars, aphids, and thrips threaten dahlias because they can eat the petals, leave slime trials, leave tattered petals, etc. Dahlias can also become infected with the following diseases: Sclerotinia disease, fungal diseases, mildew, Botrytis, Crown Gall, etc. If dahlias do become infected with these they can wilt, have spots on the leaves, the leaves can get irregular coloring/ patterning, etc.

A study on Dahlia pinnata showed that this D. pinnata was able to accumulate high levels of arsenic from the soil through its roots.

Cultivation
Used as an ornamental plant, and was cultivated by the Aztecs before the discovery of America, and was introduced to Spain in 1798. Modern dahlias are often the product of hybridisation between D. pinnata and D. coccinea. As cutflowers, dahlias have a long lifespan.

Besides being used for their outside appearance, dahlias tend to be used for their medicinal properties as well. According to Glenn Ross Whitley, this plant's roots contain some inulin and the tubers have "antibiotic compounds concentrated in [their] skin". This apparently contributed to the status of the plant as an "important root crop and medicinal plant among the pre-Columbian Indians of central Mexico, Yucatan and Guatemala".

Dahlias prefer rich soil (pH level estimated at around 6.5–7.5) with enough organic matter. The roots must be kept moist since they are very shallow rooted which means they usually become dry fairly frequently and quickly. They bloom around mid-summer through the beginning of winter, they are able to survive a light frost, but anything colder/harsher than that, they will not be able to keep thriving. When the harsh weather of winter is approaching, the dahlias should be dug up and stored in a safer place for them.

Many dahlia pinnata types can grow from seeds but more often they are cultivated by division of the tuberous roots or by stem cuttings. Providing the flower with some kind of plant food (mulch, growing media, nutrient food, etc.) can really make the dahlia healthier and more likely to survive.

Uses
The plant's phytostabilization properties regarding arsenic could lower the risk of the harmful carcinogen to humans.

Mums - Chrysanths (Chrysanthemums) by [email protected]

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Mums - Chrysanths (Chrysanthemums)

Chrysanthemums, sometimes called mums or chrysanths, are flowering plants of the genus Chrysanthemum in the family Asteraceae. They are native to East Asia and northeastern Europe. Most species originate from East Asia and the center of diversity is in China. Countless horticultural varieties and cultivars exist.

Description
The genus Chrysanthemum are perennial herbaceous flowering plants, sometimes subshrubs. The leaves are alternate, divided into leaflets and may be pinnatisect, lobed, or serrate (toothed) but rarely entire; they are connected to stalks with hairy bases.

The compound inflorescence is an array of several flower heads, or sometimes a solitary head. The head has a base covered in layers of phyllaries. The simple row of ray florets is white, yellow, or red. The disc florets are yellow. Pollen grains are approximately 34 microns.[citation needed]

Etymology
The name "chrysanthemum" is derived from the Ancient Greek: χρυσός chrysos (gold) and Ancient Greek: ἄνθεμον anthemon (flower).

Taxonomy
The genus Chrysanthemum was first formally described by Linnaeus in 1753, with 14 species, and hence bears his name (L.) as the botanical authority. The genus once included more species, but was split several decades ago into several genera, putting the economically important florist's chrysanthemums in the genus Dendranthema. The naming of these genera has been contentious, but a ruling of the International Botanical Congress in 1999 changed the defining species of the genus to Chrysanthemum indicum, restoring the florist's chrysanthemums to the genus Chrysanthemum.[citation needed]

Genera now separated from Chrysanthemum include Argyranthemum, Glebionis, Leucanthemopsis, Leucanthemum, Rhodanthemum, and Tanacetum.

Species
As of February 2020, Plants of the World Online accepted the following species:

Chrysanthemum aphrodite Kitam.
Chrysanthemum arcticum L.
Chrysanthemum argyrophyllum Ling
Chrysanthemum arisanense Hayata
Chrysanthemum chalchingolicum Grubov
Chrysanthemum chanetii H.Lév.
Chrysanthemum crassum (Kitam.) Kitam.
Chrysanthemum cuneifolium Kitam.
Chrysanthemum daucifolium Pers.
Chrysanthemum dichrum (C.Shih) H.Ohashi & Yonek.
Chrysanthemum foliaceum (G.F.Peng, C.Shih & S.Q.Zhang) J.M.Wang & Y.T.Hou
Chrysanthemum glabriusculum (W.W.Sm.) Hand.-Mazz.
Chrysanthemum horaimontanum Masam.
Chrysanthemum hypargyreum Diels
Chrysanthemum indicum L.
Chrysanthemum integrifolium Richardson
Chrysanthemum japonense (Makino) Nakai
Chrysanthemum × konoanum Makino
Chrysanthemum lavandulifolium Makino
Chrysanthemum leucanthum (Makino) Makino
Chrysanthemum longibracteatum (C.Shih, G.F.Peng & S.Y.Jin) J.M.Wang & Y.T.Hou
Chrysanthemum maximoviczii Kom.
Chrysanthemum miyatojimense Kitam.
Chrysanthemum × morifolium (Ramat.) Hemsl.
Chrysanthemum morii Hayata
Chrysanthemum naktongense Nakai
Chrysanthemum ogawae Kitam.
Chrysanthemum okiense Kitam.
Chrysanthemum oreastrum Hance
Chrysanthemum ornatum Hemsl.
Chrysanthemum parvifolium C.C.Chang
Chrysanthemum potentilloides Hand.-Mazz.
Chrysanthemum rhombifolium (Y.Ling & C.Shih) H.Ohashi & Yonek.
Chrysanthemum × rubellum Sealy
Chrysanthemum × shimotomaii Makino
Chrysanthemum sinuatum Ledeb.
Chrysanthemum vestitum (Hemsl.) Kitam.
Chrysanthemum yantaiense M.Sun & J.T.Chen
Chrysanthemum yoshinaganthum Makino
Chrysanthemum zawadskii Herbich
Chrysanthemum zhuozishanense L.Q.Zhao & Jie Yang
Former species include:

Chrysanthemum carinatum = Ismelia carinata
Chrysanthemum cinerariifolium = Tanacetum cinerariifolium
Chrysanthemum coccineum = Tanacetum coccineum
Chrysanthemum coronarium = Glebionis coronaria
Chrysanthemum frutescens = Argyranthemum frutescens
Chrysanthemum maximum = Leucanthemum maximum
Chrysanthemum pacificum = Ajania pacifica
Chrysanthemum segetum = Glebionis segetum
Ecology
Chrysanthemums start blooming in early autumn. This is also known as the favorite flower for the month of November.

History
Chrysanthemums (Chinese: 菊花; pinyin: Júhuā) were first cultivated in China as a flowering herb as far back as the 15th century BCE. Over 500 cultivars had been recorded by 1630. By 2014 it was estimated that there were over 20,000 cultivars in the world and about 7,000 cultivars in China.[14] The plant is renowned as one of the Four Gentlemen (四君子) in Chinese and East Asian Art. The plant is particularly significant during the Double Ninth Festival.

Chrysanthemum cultivation in Japan began during the Nara and Heian periods (early 8th to late 12th centuries), and gained popularity in the Edo period (early 17th to late 19th century). Many flower shapes, colours, and varieties were created. The way the flowers were grown and shaped also developed, and chrysanthemum culture flourished. Various cultivars of chrysanthemums created in the Edo period were characterized by a remarkable variety of flower shapes, and were exported to China from the end of the Edo period, changing the way Chinese chrysanthemum cultivars were grown and their popularity. In addition, from the Meiji period (late 19th to early 20th century), many cultivars with flowers over 20 cm (7.87 in) in diameter, called the Ogiku (lit., great chrysanthemum) style were created, which influenced the subsequent trend of chrysanthemums. The Imperial Seal of Japan is a chrysanthemum and the institution of the monarchy is also called the Chrysanthemum Throne. A number of festivals and shows take place throughout Japan in autumn when the flowers bloom. Chrysanthemum Day (菊の節句, Kiku no Sekku) is one of the five ancient sacred festivals. It is celebrated on the 9th day of the 9th month. It was started in 910, when the imperial court held its first chrysanthemum show.

Chrysanthemums entered American horticulture in 1798 when Colonel John Stevens imported a cultivated variety known as 'Dark Purple' from England. The introduction was part of an effort to grow attractions within Elysian Fields in Hoboken, New Jersey.

Modern cultivated chrysanthemums are showier than their wild relatives. Many horticultural specimens have been bred to bear many rows of ray florets in a great variety of colors. The flower heads occur in various forms, and can be daisy-like or decorative, like pompons or buttons. This genus contains many hybrids and thousands of cultivars developed for horticultural purposes. In addition to the traditional yellow, other colors are available, such as white, purple, and red. The most important hybrid is Chrysanthemum × morifolium (syn. C. × grandiflorum), derived primarily from C. indicum, but also involving other species.[citation needed]

Over 140 cultivars of chrysanthemum have gained the Royal Horticultural Society's Award of Garden Merit (confirmed 2017).

In Japan, a form of bonsai chrysanthemum was developed over the centuries. The cultivated flower has a lifespan of about 5 years and can be kept in miniature size. Another method is to use pieces of dead wood and the flower grows over the back along the wood to give the illusion from the front that the miniature tree blooms.

Culinary uses
Yellow or white chrysanthemum flowers of the species C. morifolium are boiled to make a tea in some parts of East Asia. The resulting beverage is known simply as chrysanthemum tea (菊 花 茶, pinyin: júhuā chá, in Chinese). In Korea, a rice wine flavored with chrysanthemum flowers is called gukhwaju (국화주).

Chrysanthemum leaves are steamed or boiled and used as greens, especially in Chinese cuisine. The flowers may be added to dishes such as mixian in broth, or thick snakemeat soup (蛇羹) to enhance the aroma. They are commonly used in hot pot and stir fries. Japanese cuisine sashimi uses small chrysanthemums as garnish.

Insecticidal uses
Pyrethrum (Chrysanthemum [or Tanacetum] cinerariaefolium) is economically important as a natural source of insecticide. The flowers are pulverized, and the active components, called pyrethrins, which occur in the achenes, are extracted and sold in the form of an oleoresin. This is applied as a suspension in water or oil, or as a powder. Pyrethrins attack the nervous systems of all insects, and inhibit female mosquitoes from biting. In sublethal doses, they have an insect repellent effect.[citation needed] They are harmful to fish, but are far less toxic to mammals and birds than many synthetic insecticides. They are not persistent, being biodegradable, and also decompose easily on exposure to light. Pyrethroids such as permethrin are synthetic insecticides based on natural pyrethrum. Despite this, chrysanthemum leaves are still a major host for destructive pests, such as leafminer flies including L. trifolii.

Environmental uses
Chrysanthemum plants have been shown to reduce indoor air pollution by the NASA Clean Air Study

Cultural significance and symbolism
In some European countries (e.g., France, Belgium, Italy, Spain, Poland, Hungary, Croatia), incurve chrysanthemums symbolize death and are used only for funerals or on graves, while other types carry no such symbolism;[citation needed] similarly, in China, Japan, and Korea of East Asia, white chrysanthemums symbolize adversity, lamentation, and/or grief. In some other countries, they represent honesty. In the United States, the flower is usually regarded as positive and cheerful, with New Orleans as a notable exception.

In the Victorian language of flowers, the chrysanthemum had several meanings. The Chinese chrysanthemum meant cheerfulness, whereas the red chrysanthemum stood for "I Love", while the yellow chrysanthemum symbolized slighted love. The chrysanthemum is also the flower of November.

The chrysanthemum is the city flower of Beijing and Kaifeng. The tradition of cultivating different varieties of chrysanthemums stretches back 1600 years, and the scale reached a phenomenal level during the Song dynasty until its loss to the Jürchens in 1126. The city has held the Kaifeng Chrysanthemum Cultural Festival since 1983 (renamed China Kaifeng Chrysanthemum Cultural Festival in 1994). The event is the largest chrysanthemum festival in China; it has been a yearly feature since, taking place between 18 October and 18 November every year.
The chrysanthemum is one of the "Four Gentlemen" (四君子) of China (the others being the plum blossom, the orchid, and bamboo). The chrysanthemum is said to have been favored by Tao Qian, an influential Chinese poet, and is symbolic of nobility. It is also one of the four symbolic seasonal flowers.
A chrysanthemum festival is held each year in Tongxiang, near Hangzhou, China.
Chrysanthemums are the topic in hundreds of poems of China.
The "golden flower" referred to in the 2006 movie Curse of the Golden Flower is a chrysanthemum.
"Chrysanthemum Gate" (jú huā mén 菊花门), often abbreviated as Chrysanthemum (菊花), is taboo slang meaning "anus" (with sexual connotations).
An ancient Chinese city (Xiaolan Town of Zhongshan City) was named Ju-Xian, meaning "chrysanthemum city".
The plant is particularly significant during the Chinese Double Ninth Festival.
In Chinese culture, the chrysanthemum is a symbol of autumn and the flower of the ninth moon. People even drank chrysanthemum wine on the ninth day of the ninth lunar month to prolong their lives during the Han dynasty. It is a symbol of longevity because of its health-giving properties. Because of all of this, the flower was often worn on funeral attire.
Pharmacopoeia of the People's Republic of China listed two kinds of chrysanthemum for medical use, Yejuhua and Juhua. Historically Yejuhua is said to treat carbuncle, furuncle, conjunctivitis, headache, and vertigo. Juhua is said to treat cold, headache, vertigo, and conjunctivitis.
Japan

Chrysanthemums and fences.
Chrysanthemums first arrived in Japan by way of China in the 5th century. The chrysanthemum has been used as a theme of waka (Japanese traditional poetry) since around the 10th century in the Heian period, and Kokin Wakashū is the most famous of them. In the 12th century, during the Kamakura period, when the Retired Emperor Go-Toba adopted it as the mon (family crest) of the Imperial family, it became a flower that symbolized autumn in Japan. During the Edo period from the 17th century to the 19th century, due to the development of economy and culture, the cultivation of chrysanthemums, cherry blossoms, Japanese iris, morning glory, etc. became popular, many cultivars were created and many chrysanthemum exhibitions were held. From the Meiji period in the latter half of the 19th century, due to the growing importance of the chrysanthemum, which symbolized the Imperial family, the creation of ogiku style cultivars with a diameter of 20 cm or more became popular.

In the present day, each autumn there are chrysanthemum exhibitions at the Shinjuku Gyo-en, Meiji Shrine and Yasukuni Shrine in Tokyo. The Yasukuni Shrine, formerly a state-endowed shrine (官国弊社, kankokuheisha) has adopted the chrysanthemum crest. Culinary-grade chrysanthemums are used to decorate food, and they remain a common motif for traditional Japanese arts like porcelain, lacquerware and kimono.

Chrysanthemum growing is still practised actively as a hobby by many Japanese people who enter prize plants in contests. Chrysanthemum "dolls", often depicting fictional characters from both traditional sources like kabuki and contemporary sources like Disney, are displayed throughout the fall months, and the city of Nihonmatsu hosts the "Nihonmatsu Chrysanthemum Dolls Exhibition" every autumn in historical ruin of Nihonmatsu Castle. They are also grown into chrysanthemum bonsai forms.

In Japan, the chrysanthemum is a symbol of the Emperor and the Imperial family. In particular, a "chrysanthemum crest" (菊花紋章, kikukamonshō or kikkamonshō), i.e. a mon of chrysanthemum blossom design, indicates a link to the Emperor; there are more than 150 patterns of this design. Notable uses of and reference to the Imperial chrysanthemum include:
The Imperial Seal of Japan is used by members of the Japanese imperial family. In 1869, a two-layered, 16-petal design was designated as the symbol of the emperor. Princes used a simpler, single-layer pattern.
The Chrysanthemum Throne is the name given to the position of Japanese Emperor and the throne.
The Supreme Order of the Chrysanthemum is a Japanese honor awarded by the emperor on the advice of the Japanese government.
In Imperial Japan, small arms were required to be stamped with the imperial chrysanthemum, as they were considered the personal property of the emperor.
The Nagoya Castle Chrysanthemum Competition started after the end of the Pacific War. The event at the castle has become a tradition for the city. With three categories, it is one of the largest events of its kind in the region by both scale and content. The first category is the exhibition of cultivated flowers. The second category is for bonsai flowers, which are combined with dead pieces of wood to give the illusion of miniature trees. The third category is the creation of miniature landscapes.

Asia
Korea
Korea has a number of flower shows that exhibit the chrysanthemum, such as the Masan Gagopa Chrysanthemum Festival.

West Asia
Iran
In Iran, chrysanthemums are associated with the Zoroastrian spiritual being Ashi Vanghuhi (lit. 'good blessings, rewards'), a female Yazad (angel) presiding over blessings.

Israel
In 2017, Israel named a new fast-growing Chrysanthemum flower after Narendra Modi, the Prime Minister of India, in a special gesture to mark the first visit of an Indian prime minister to the Jewish nation.

Oceania
Australia
In Australia, on Mother's Day, which falls in May when the flower is in season, people traditionally wear a white chrysanthemum, or a similar white flower to honour their mothers. Chrysanthemums are often given as Mother's Day presents.

North America
United States
On 5 and 6 November 1883, in Philadelphia, the Pennsylvania Horticultural Society (PHS), at the request of the Florists and Growers Society, held its first Chrysanthemum Show in Horticultural Hall. This would be the first of several chrysanthemum events presented by PHS to the public.
The founding of the chrysanthemum industry dates back to 1884, when the Enomoto brothers of Redwood City, California grew the first chrysanthemums cultivated in America.
In 1913, Sadakasu Enomoto (of San Mateo County) astounded the flower world by successfully shipping a carload of Turner chrysanthemums to New Orleans for the All Saints Day Celebration.
The chrysanthemum was recognized as the official flower of the city of Chicago by Mayor Richard J. Daley in 1966.
The chrysanthemum is the official flower of the city of Salinas, California.
The chrysanthemum is the official flower of several fraternities and sororities including Chi Phi, Phi Kappa Sigma, Phi Mu Alpha Sinfonia, Lambda Kappa Sigma, Sigma Alpha and Triangle Fraternity.

Europe
Italy
Italian composer Giacomo Puccini wrote Crisantemi (1890), a movement for string quartet, in memory of his friend Amedeo di Savoia Duca d'Aosta. In Italy (and other European countries) the chrysanthemum is the flower that people traditionally bring to their deceased loved ones at the cemetery and is generally associated with mourning. A probable reason for this is the fact that the plant flowers between the end of October and the beginning of November, coinciding with the Day of the Dead (2 November).

Poland
Chrysanthemums are placed on graves to honor the dead during All Saints' Day and All Souls' Day in Poland.

United Kingdom
The UK National Collection of hardy chrysanthemums is at Hill Close Gardens near Warwick.

Dimorphotheca by [email protected]

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Dimorphotheca

Dimorphotheca is a genus of plants in the family Asteraceae, native to southern Africa. is one of eight genera of the Calenduleae, with a centre of diversity in Southern Africa. Species are native to Botswana, Eswatini, Lesotho, Namibia, South Africa, and Zimbabwe. Some species can hybridize with Osteospermum, and crosses are sold as cultivated ornamentals. The name "Dimorphotheca" comes from the Greek "Dis" "Morphe" and "Theka", meaning "two shaped receptacle", referring to the dimorphic cypselae, a trait inherent to members of the Calenduleae. Plants of this genus usually have bisexual flowers.

Species
Dimorphotheca acutifolia
Dimorphotheca barberae
Dimorphotheca caulescens
Dimorphotheca chrysanthemifolia
Dimorphotheca cuneata
Dimorphotheca dregei
Dimorphotheca ecklonis
Dimorphotheca fruticosa
Dimorphotheca jucunda
Dimorphotheca montana
Dimorphotheca nudicaulis
Dimorphotheca pluvialis ("ox-eye daisy", "Cape daisy", "rain daisy")
Dimorphotheca polyptera
Dimorphotheca pulvinalis
Dimorphotheca sinuata ("Cape marigold", "African daisy", "star of the veldt")
Dimorphotheca spectabilis
Dimorphotheca tragus
Dimorphotheca turicensis
Dimorphotheca venusta
Dimorphotheca walliana
Dimorphotheca zeyheri

Daisybushes - African Daisies - African Daisy - South African Daisy - Cape Daisy - Blue-Eyed Daisy (Osteospermum) by [email protected]

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Daisybushes - African Daisies - African Daisy - South African Daisy - Cape Daisy - Blue-Eyed Daisy (Osteospermum)

Osteospermum is a genus of flowering plants belonging to the Calenduleae, one of the smaller tribes of the sunflower/daisy family Asteraceae. They are known as the daisybushes or African daisies. Its species have been given several common names, including African daisy, South African daisy, Cape daisy and blue-eyed daisy.

Description
Their alternate (rarely opposite) leaves are green, but some variegated forms exist. The leaf form is lanceolate. The leaf margin is entire, but hardy types are toothed.

The daisy-like composite flower consists of disc florets and ray florets, growing singly at the end of branches or sometimes in inflorescences of terminal corymbose cymes. The disc florets are pseudo-bisexual and come in several colors such as blue, yellow and purple. The hardy types usually show a dark blue center in the disc until the yellow pollen is shed. The ray florets are female and are found diverse colors such as white, cream, pink, purple, mauve to yellow. Some cultivars have "spooned" petals such as "Pink Whirls". Many species flower a second time late summer, stimulated by the cooler night temperatures. Hardy types show profuse flowering in the spring, but they do not get a second flush of flowers.

Taxonomy
The genus Osteospermum was named by Carl Linnaeus in 1753. The scientific name is derived from the Greek osteon (bone) and Latin spermum (seed). Plants of the World Online and the South African National Biodiversity Institute accept a broad definition of the genus, which is a sister taxon to Dimorphotheca. Other authorities treat its generic synonyms (Chrysanthemoides, Oligocarpus, etc.) as separate genera.

Species
74 species are accepted:

Osteospermum acanthospermum (DC.) Norl.
Osteospermum aciphyllum DC.
Osteospermum afromontanum Norl.
Osteospermum amplectens (Harv.) Norl.
Osteospermum angolense Norl.
Osteospermum apterum (B.Nord.) J.C.Manning & Goldblatt
Osteospermum armatum Norl.
Osteospermum asperulum (DC.) Norl.
Osteospermum attenuatum Hilliard & B.L.Burtt
Osteospermum auriculatum (S.Moore) Norl.
Osteospermum australe B.Nord.
Osteospermum bidens Thunb.
Osteospermum bolusii (Compton) Norl.
Osteospermum breviradiatum Norl. — Lemoenboegoe
Osteospermum burttianum B.Nord.
Osteospermum calcicola (J.C.Manning & Goldblatt) J.C.Manning & Goldblatt
Osteospermum calendulaceum L.f. — Stinking Roger
Osteospermum ciliatum P.J.Bergius
Osteospermum connatum DC.
Osteospermum corymbosum L.
Osteospermum crassifolium (O.Hoffm.) Norl.
Osteospermum dentatum Burm.f.
Osteospermum elsieae Norl.
Osteospermum grandidentatum DC. — Yellow trailing daisy
Osteospermum grandiflorum DC.
Osteospermum hafstroemii Norl.
Osteospermum herbaceum L.f.
Osteospermum hirsutum Thunb.
Osteospermum hispidum Harv.
Osteospermum hyoseroides (DC.) Norl.
Osteospermum ilicifolium L.
Osteospermum imbricatum L.
Osteospermum incanum Burm.f.
Osteospermum junceum P.J.Bergius
Osteospermum karrooicum (Bolus) Norl.
Osteospermum lanceolatum DC.
Osteospermum leptolobum (Harv.) Norl.
Osteospermum microcarpum (Harv.) Norl.
Osteospermum microphyllum DC.
Osteospermum moniliferum L.
Osteospermum monocephalum (Oliv. & Hiern) Norl.
Osteospermum monstrosum (Burm.f.) J.C.Manning & Goldblatt
Osteospermum montanum Klatt
Osteospermum muricatum E.Mey. ex DC.
Osteospermum namibense Swanepoel
Osteospermum nervosum (Hutch.) Norl.
Osteospermum nordenstamii J.C.Manning & Goldblatt
Osteospermum norlindhianum J.C.Manning & Goldblatt
Osteospermum nyikense Norl.
Osteospermum oppositifolium (Aiton) Norl.
Osteospermum pinnatilobatum Norl.
Osteospermum pinnatum (Thunb.) Norl.
Osteospermum polycephalum (DC.) Norl.
Osteospermum polygaloides L.
Osteospermum potbergense A.R.Wood & B.Nord.
Osteospermum pterigoideum Klatt
Osteospermum pyrifolium Norl.
Osteospermum rigidum Aiton
Osteospermum rosulatum Norl.
Osteospermum rotundifolium (DC.) Norl.
Osteospermum sanctae-helenae Norl.
Osteospermum scariosum DC.
Osteospermum sinuatum (DC.) Norl.
Osteospermum spathulatum (DC.) Norl.
Osteospermum spinescens Thunb.
Osteospermum spinigerum (Norl.) Norl.
Osteospermum spinosum L.
Osteospermum striatum Burtt Davy
Osteospermum subulatum DC.
Osteospermum thodei Markötter
Osteospermum tomentosum (L.f.) B.Nord.
Osteospermum triquetrum L.f.
Osteospermum vaillantii (Decne.) Norl.
Osteospermum volkensii (O.Hoffm.) Norl.
Formerly placed here
Dimorphotheca barberae — synonym Osteospermum barberae (Harv.) Norl.)
Dimorphotheca ecklonis — Cape marguerite, blue-and-white daisybush (synonym Osteospermum ecklonis (DC.) Norl.)
Dimorphotheca fruticosa — Trailing African daisy, shrubby daisybush (synonym Osteospermum fruticosum (L.) Norl.)
Dimorphotheca jucunda (E.Phillips) Norl. — South African daisy (synonym Osteospermum jucundum E.Phillips)
Distribution
There are about 70 species native to southern and eastern Africa and the Arabian peninsula.

Cultivation
Osteospermum are popular in cultivation, where they are frequently used in summer bedding schemes in parks and gardens. Numerous hybrids and cultivars have been grown with a wide range of tropical colors. Yellow cultivars tend to have a yellow center (sometimes off-white).

Plants prefer a warm and sunny position and rich soil, although they tolerate poor soil, salt or drought well. Modern cultivars flower continuously when watered and fertilised well, and dead-heading is not necessary, because they do not set seed easily. If planted in a container, soil should be prevented from drying out completely. If they do, the plants will go into "sleep mode" and survive the period of drought, but they will abort their flower buds and not easily come back into flower. Moreover, roots are relatively susceptible to rotting if watered too profusely after the dry period.

Cultivars
Most widely sold cultivars are grown as annuals, are mainly hybrids of O. jucundum, O. ecklonis and O. grandiflorum and can be hardy to -2 °C (30 °F). If hardy, they can be grown as perennials or as shrubs.

Cultivars (those marked agm have gained the Royal Horticultural Society's Award of Garden Merit):

'Acapulco'
'African Queen'
'Apricot'
'Biera'
'Big Pink'
‘Blackthorn Seedling’ agm
'Bodegas Pink'
'Buttermilk' agm[11]
'Chris Brickell'
'Duet'
'Giles Gilbey'
'Hopleys’ agm
'Ice White'
'Langtrees agm
’Lady Leitrim’ agm
'Lilac Spoon'
'Marbella'
'Merriments Joy'
'Nairobi Purple'
O. jucundum agm
'Passion Mix'
'Pink'
'Pink Beauty'
'Pink Whirls' agm
'Silver Sparkler' agm
'Soprano'
'Starshine'
'Springstar Gemma'
'Sunkist'
'Weetwood'
'White Pim' agm
'White Spoon'
'White Whirls'
'Whirlygig'

Dimorphotheca by [email protected]

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Dimorphotheca

Dimorphotheca is a genus of plants in the family Asteraceae, native to southern Africa. is one of eight genera of the Calenduleae, with a centre of diversity in Southern Africa. Species are native to Botswana, Eswatini, Lesotho, Namibia, South Africa, and Zimbabwe. Some species can hybridize with Osteospermum, and crosses are sold as cultivated ornamentals. The name "Dimorphotheca" comes from the Greek "Dis" "Morphe" and "Theka", meaning "two shaped receptacle", referring to the dimorphic cypselae, a trait inherent to members of the Calenduleae. Plants of this genus usually have bisexual flowers.

Species
Dimorphotheca acutifolia
Dimorphotheca barberae
Dimorphotheca caulescens
Dimorphotheca chrysanthemifolia
Dimorphotheca cuneata
Dimorphotheca dregei
Dimorphotheca ecklonis
Dimorphotheca fruticosa
Dimorphotheca jucunda
Dimorphotheca montana
Dimorphotheca nudicaulis
Dimorphotheca pluvialis ("ox-eye daisy", "Cape daisy", "rain daisy")
Dimorphotheca polyptera
Dimorphotheca pulvinalis
Dimorphotheca sinuata ("Cape marigold", "African daisy", "star of the veldt")
Dimorphotheca spectabilis
Dimorphotheca tragus
Dimorphotheca turicensis
Dimorphotheca venusta
Dimorphotheca walliana
Dimorphotheca zeyheri

Cape Marguerite - African Daisy - Van Staden's River Daisy - Sundays River Daisy - White Daisy Bush - Blue & White Daisy Bush - Star Of The Veldt (Dimorphotheca Ecklonis) by [email protected]

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Cape Marguerite - African Daisy - Van Staden's River Daisy - Sundays River Daisy - White Daisy Bush - Blue & White Daisy Bush - Star Of The Veldt (Dimorphotheca Ecklonis)

Dimorphotheca is a genus of plants in the family Asteraceae, native to southern Africa. is one of eight genera of the Calenduleae, with a centre of diversity in Southern Africa. Species are native to Botswana, Eswatini, Lesotho, Namibia, South Africa, and Zimbabwe. Some species can hybridize with Osteospermum, and crosses are sold as cultivated ornamentals. The name "Dimorphotheca" comes from the Greek "Dis" "Morphe" and "Theka", meaning "two shaped receptacle", referring to the dimorphic cypselae, a trait inherent to members of the Calenduleae. Plants of this genus usually have bisexual flowers.

Species
Dimorphotheca acutifolia
Dimorphotheca barberae
Dimorphotheca caulescens
Dimorphotheca chrysanthemifolia
Dimorphotheca cuneata
Dimorphotheca dregei
Dimorphotheca ecklonis
Dimorphotheca fruticosa
Dimorphotheca jucunda
Dimorphotheca montana
Dimorphotheca nudicaulis
Dimorphotheca pluvialis ("ox-eye daisy", "Cape daisy", "rain daisy")
Dimorphotheca polyptera
Dimorphotheca pulvinalis
Dimorphotheca sinuata ("Cape marigold", "African daisy", "star of the veldt")
Dimorphotheca spectabilis
Dimorphotheca tragus
Dimorphotheca turicensis
Dimorphotheca venusta
Dimorphotheca walliana
Dimorphotheca zeyheri

Dimorphotheca ecklonis, also known as Cape marguerite, African daisy, Van Staden's river daisy, Sundays river daisy, white daisy bush, blue-and-white daisy bush, star of the veldt is an ornamental plant that is native to South Africa. It is now regarded as a weed in parts of Australia, particularly Victoria and Western Australia.

Features
It is an evergreen, perennial dwarf shrub with the stature heights of 25 to 50 centimeters. The leaves measure 5 to 10 × 1 to 4 inches and are glandular fluffy, sessile, elliptical, slightly succulent, narrow obovate and entire or serrated. The heads are on 15 to 20 centimeters long stems and have a diameter of 5 to 8 centimeters. The bracts are 13 to 16 millimeters long and glandular. The rays are white above and reddish blue below. The disc is dark blue or purple. The fruit surface is net-wrinkled.

The flowering period is from April to September in the northern hemisphere. The plant is said to contain hydrocyanic acid, making it poisonous to livestock.

Distribution
It is native to the Eastern Cape in Uitenhage and Humansdorp, in South Africa, and is found on wet grass and in river beds at altitudes up to 300 meters.

Cultivation
It is widely used as an ornamental plant in pots, summer borders and balcony boxes. It is cultivated one-year old. There are numerous hybrids and varieties, including upright, up to 1.5 meters high and half-low. This species has been in culture since about 1920. It thrives best in full sun, in poor, sandy soil.

Dimorphotheca by [email protected]

© [email protected], all rights reserved.

Dimorphotheca

Dimorphotheca is a genus of plants in the family Asteraceae, native to southern Africa. is one of eight genera of the Calenduleae, with a centre of diversity in Southern Africa. Species are native to Botswana, Eswatini, Lesotho, Namibia, South Africa, and Zimbabwe. Some species can hybridize with Osteospermum, and crosses are sold as cultivated ornamentals. The name "Dimorphotheca" comes from the Greek "Dis" "Morphe" and "Theka", meaning "two shaped receptacle", referring to the dimorphic cypselae, a trait inherent to members of the Calenduleae. Plants of this genus usually have bisexual flowers.

Species
Dimorphotheca acutifolia
Dimorphotheca barberae
Dimorphotheca caulescens
Dimorphotheca chrysanthemifolia
Dimorphotheca cuneata
Dimorphotheca dregei
Dimorphotheca ecklonis
Dimorphotheca fruticosa
Dimorphotheca jucunda
Dimorphotheca montana
Dimorphotheca nudicaulis
Dimorphotheca pluvialis ("ox-eye daisy", "Cape daisy", "rain daisy")
Dimorphotheca polyptera
Dimorphotheca pulvinalis
Dimorphotheca sinuata ("Cape marigold", "African daisy", "star of the veldt")
Dimorphotheca spectabilis
Dimorphotheca tragus
Dimorphotheca turicensis
Dimorphotheca venusta
Dimorphotheca walliana
Dimorphotheca zeyheri

China Aster - Annual Aster (Callistephus Chinensis) by [email protected]

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China Aster - Annual Aster (Callistephus Chinensis)

Callistephus is a monotypic genus of flowering plants in the aster family, Asteraceae, containing the single species Callistephus chinensis. Its common names include China aster and annual aster. It is native to China and Korea and it is cultivated worldwide as an ornamental plant in cottage gardens and as a cut flower.

Description
This is an annual or biennial plant with one erect, mostly unbranched stem growing 20–100 cm (7.9–39.4 in) tall. The alternately arranged leaves vary in shape or size. The basal leaves usually fall away before flowering. Leaves around the middle of the stem are a few centimeters long and are borne on winged petioles. The blades have serrated edges. Leaves toward the top may have smooth edges. The large solitary flower head grows at the top of the stem and sometimes on branches. The head is lined with layers of phyllaries, those in the outer layer large and leaflike, measuring up to 3 centimeters long. The inner layer can be purple-tinged. The head contains one or two rings of ray florets, most often in shades of reddish purple. The flowering period is from August to October in the northern hemisphere.

Cultivation
This species has been grown and bred extensively in cultivation. Cultivars are available with flowers of many colors, varying heights, and single and double heads. In the wild, they are purple in color. There are several cultivars with flowers of different colors. Cultivars with flowers of other colors were obtained in particular by hybridization. Cultivated varieties can have ray florets in most any color, including red, pink, blue, purple, and white. They are up to 3.5 centimeters long. There are many yellow disc florets in the center. The fruit is a rough-textured, glandular, purple-mottled cypsela that turns gray with age. It has a pappus composed of one outer layer of reddish scales and two inner layers of white bristles.

It has been in cultivation in Europe since 1728 at the latest. In China, it has been cultivated for 2000 years.

Distribution
This plant is a naturalized species in some areas outside of its native range (Indochina, Europe, North America, Australia, New Zealand, etc.) and can be found on the edges of deciduous forests. It grows at altitudes from 300 to 2700 above mean sea level. As ornamental plant it thrives on fresh, nutrient-rich, easy-humus rich soils in warm humid air conditioning.

Pests and diseases
The plants are susceptible to fusarium wilt, a fungal disease. Some cultivars are more resistant to this than others. Pest insects can include leafhoppers, aphids, blister beetles, and the tarnished plant bug. The plant is also susceptible to mites.

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